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The Zopheridae family of
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s has grown considerably in recent years as the members of two other families have been included within its circumscription; these former families are the
Monommatidae Monommatini is a tribe of beetles known as monommatid beetles. They are in the ironclad beetle family, Zopheridae. There are about 15 genera and 300 described species in Monommatini. They are found worldwide, with the greatest diversity in Madaga ...
and the
Colydiidae Colydiinae is a subfamily of beetles, commonly known as cylindrical bark beetles. They have been treated historically as a family Colydiidae, but have been moved into the Zopheridae, where they constitute the bulk of the diversity of the newly e ...
, which are now both included in the Zopheridae as
subfamilies In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
or (in the former case) even as
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...
of
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Zopherinae Zopherinae is a subfamily of beetles, commonly known as ironclad beetles. Together with the subfamily Usechinae, they have been treated historically as a family, but have recently been joined by several additional taxa, making the Zopheridae ...
. Some authors accept up to six subfamilies here, while others merge all except the Colydiinae into the
Zopherinae Zopherinae is a subfamily of beetles, commonly known as ironclad beetles. Together with the subfamily Usechinae, they have been treated historically as a family, but have recently been joined by several additional taxa, making the Zopheridae ...
. The family has approximately 190 genera and 1700 species, which are found worldwide. A large number of members of the family feed on rotting wood or fungus associated with rotting wood, though some members of
Colydiinae Colydiinae is a subfamily of beetles, commonly known as cylindrical bark beetles. They have been treated historically as a family Colydiidae, but have been moved into the Zopheridae, where they constitute the bulk of the diversity of the newly ex ...
are predatory, or feed on living plant tissue such as roots, stems, flower stalks and fruit. The oldest fossils of the family are '' Paleoendeitoma'' (subfamily
Colydiinae Colydiinae is a subfamily of beetles, commonly known as cylindrical bark beetles. They have been treated historically as a family Colydiidae, but have been moved into the Zopheridae, where they constitute the bulk of the diversity of the newly ex ...
, tribe Synchitini) and '' Cretomysteria'' from the early Late Cretaceous (
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
)
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...
from Myanmar.


See also

*
List of subgroups of the order Coleoptera This article scientific classification, classifies the subgroups of the order Coleoptera (beetles) down to the level of family (biology), families, following the system in "Family-group names in Coleoptera (Insecta)", Bouchard, et al. (2011), with ...


Genera

These 89 genera belong to the family Zopheridae: * '' Ablabus'' Broun, 1880 g * '' Acolobicus'' Sharp, 1894 i c g b * '' Acropis'' Burmeister, 1840 g * '' Anarmostes'' Pascoe, 1860 g * '' Anchomma'' Leconte, 1858 g * '' Antilissus'' Sharp, 1879 i c g * ''
Aspathines Aspathines ( peo, wiktionary:𐎠𐎿𐎱𐎨𐎴𐎠, 𐎠𐎿𐎱𐎨𐎴𐎠 ; grc, wiktionary:Ἀσπαθίνης, Ἀσπαθίνης ) (born and died sometime between 550 BC and 450BC) was a senior official under Darius the Great and Xerxes ...
'' Champion, 1888 i c g b * '' Aulonium'' Erichson, 1845 i c g b * '' Bitoma'' Herbst, 1793 i c g b * '' Catolaemus'' Sharp, 1894 g * '' Cerchanotus'' Erichson, 1845 g * '' Chorasus'' Sharp, 1882 g * ''
Cicones The Cicones (; ) or Ciconians were a Homeric ThracianHerodotus, ''The Histories'' (Penguin Classics), edd. John M. Marincola and Aubery de Selincourt, 2003, p. 452 (I10): "The Thracian tribes lying along his route were the Paeti, Cicones, Bisto ...
'' Curtis, 1827 g * '' Colobicones'' Grouvelle, 1918 g * '' Colobicus'' Latreille, 1807 i c g b * '' Colydium'' Fabricius, 1792 i c g b * '' Colydodes'' Motschulsky, 1855 g * '' Corticus'' Germar, 1824 g * '' Coxelus'' Dejean, 1821 i c g b * '' Denophoelus'' Stephan, 1989 i c g b * '' Diodesma'' Latreille, 1829 g * '' Diplagia'' Reitter, 1882 g * '' Dryptops'' Broun, 1882 g * '' Endeitoma'' Sharp, 1894 i c g b * '' Endocoxelus'' Waterhouse, 1876 g * '' Endophloeus'' Dejean, 1834 g * '' Enhypnon'' Carter, 1919 g * '' Ethelema'' Pascoe, 1860 g * '' Eucicones'' Sharp, 1894 i c g b * '' Eudesma'' LeConte, 1863 i c g b * '' Glyphocryptus'' Sharp, 1885 g * '' Helioctamenus'' Schaufuss, 1882 g * '' Hyporhagus'' Thomson, 1860 i c g b * '' Hyporrhagus'' Thomson, 1860 g * '' Inscutomonomma'' Pic, 1951 g * '' Langelandia'' Aubé, 1842 g * '' Lasconotus'' Erichson, 1845 i c g b * '' Lemnis'' Harold, 1868 g * '' Lobogestoria'' Reitter, 1878 i c g b * '' Lyreus'' Aubé, 1861 i c g b * '' Mecedanum'' Erichson, 1845 g * '' Megataphrus'' Casey, 1890 i c g b * '' Microprius'' Fairmaire, 1868 i c g b * '' Microsicus'' Sharp, 1894 i c g * '' Monoedus'' Horn, 1882 i c g b * '' Monomma'' Klug, 1833 g * '' Myrmechixenus'' g * '' Namunaria'' Reitter, 1882 i c g b * '' Nematidium'' Erichson, 1845 i c g b * '' Neotrichus'' Sharp, 1886 i c g * '' Noserinus'' Casey, 1907 g * '' Noserus'' g * '' Nosoderma'' Solier, 1841 g * '' Nosodomodes'' Reitter, 1922 g * '' Orthocerus'' g * '' Paha'' Dajoz, 1984 i c g b * '' Phellopsis'' LeConte, 1862 i c g b * '' Phloeodalis'' Erichson, 1845 g * '' Phloeodes'' LeConte, 1862 i c g b * '' Phloeonemus'' Erichson, 1845 i c g b * '' Phormesa'' Pascoe, 1863 g * '' Phreatus'' Pascoe, 1863 g * '' Priolomus'' Erichson, 1845 g * '' Pristoderus'' Hope, 1840 g * '' Prosteca'' Wollaston, 1860 g * '' Prostheca'' Wollaston, 1860 g * '' Pseudaulonium'' Reitter, 1877 g * '' Pseudendestes'' Lawrence, 1980 g * '' Pseudocorticus'' Hinton, 1935 i c g b * '' Pycnomerus'' Erichson, 1842 i c g b * '' Rechodes'' Erichson, 1845 g * '' Recyntus'' Broun, 1882 g * '' Rhagodera'' Mannerheim, 1843 i c g b * '' Rhagoderidea'' Wickham, 1914 g * '' Rhizonium'' Sharp, 1876 g * '' Sesaspis'' b * '' Spinhyporhagus'' Freude, 1984 i c g * '' Stephaniolus'' Ivie, Slipinsky and Wegrzynowicz, 2001 i c g * '' Synchita'' Hellwig, 1792 i c g b * '' Tarphiablabus'' Slipinski & Lawrence, 1997 g * '' Tarphiosoma'' Wollaston, 1862 g * '' Tarphius'' Erichson, 1845 g * '' Todimopsis'' Slipinski & Lawrence, 1997 g * '' Trigonophloeus'' Dajoz, 1980 g * '' Usechimorpha'' Blaisdell, 1929 i c g b * '' Usechus'' Motschulsky, 1845 i c g b * '' Vitiacus'' Broun, 1893 g * '' Xylolaemus'' Dejean, 1835 g * ''
Zopherus ''Zopherus'' is a genus of beetles comprising 19 species. They live in the Americas and are adapted to wood-boring. Distribution Species of ''Zopherus'' only live in the Americas, where they are distributed from Venezuela to the southern Uni ...
'' Laporte, 1840 i c g b Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net


References

{{Taxonbar, from=Q1057389 Beetle families Tenebrionoidea