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''Zephyrarchaea marki'', the Cape Le Grand assassin spider, is a species of
spider Spiders (order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species d ...
in the family
Archaeidae Archaeidae, also known as assassin spiders and pelican spiders, is a spider family with about ninety described species in five genera. It contains small spiders, ranging from long, that prey exclusively on other spiders. They are unusual in that ...
, commonly known as the assassin spiders. Known only from
Cape Le Grand National Park Cape Le Grand National Park is a national park in Western Australia, south-east of Perth and east of Esperance. The park covers an area of The area is an ancient landscape which has been above sea level for well over 200 million years and ...
in Western Australia, the species was first described by
Michael G. Rix Michael Gordon Rix is an Australian arachnologist, whose publications mainly concern spiders. , he was Principal Curator of Arachnology and Research Fellow in the Biodiversity and Geosciences Program at the Queensland Museum. He has held numero ...
and
Mark Harvey Mark Harvey (born 11 June 1965) is a former Australian rules football player and coach. He played over 200 games during fourteen seasons with the Essendon Football Club, winning three premierships, and was senior coach of Fremantle from 2007 ...
in 2012. It is named after Mark Wojcieszek, who helped collect the initial specimens of this species. ''Z. marki'' is a small species of spider, with a total length of in adult males. In adult males, the cephalothorax is dark reddish-brown and the abdomen is mottled grey-brown and beige. The legs are tan brown with darker ring-like markings. The appearance of the female is unknown. The species is known to inhabit elevated leaf litter in a dense coastal thickets of ''
Banksia speciosa ''Banksia speciosa'', commonly known as the showy banksia, is a species of large shrub or small tree in the family Proteaceae. It is found on the south coast of Western Australia between Hopetoun (33°57′ S) and the Great Australian Bi ...
''. It has not yet been evaluated and assigned a conservation status by the International Union for Conservation of Nature. However, the species is endemic to a very small range and its only known population may be threatened by fire, dieback disease affecting ''Banksia'', and climate change.


Taxonomy

''Zephyrarchaea marki'' was described by the arachnologists 
Michael G. Rix Michael Gordon Rix is an Australian arachnologist, whose publications mainly concern spiders. , he was Principal Curator of Arachnology and Research Fellow in the Biodiversity and Geosciences Program at the Queensland Museum. He has held numero ...
and
Mark Harvey Mark Harvey (born 11 June 1965) is a former Australian rules football player and coach. He played over 200 games during fourteen seasons with the Essendon Football Club, winning three premierships, and was senior coach of Fremantle from 2007 ...
in 2012 on the basis of a male specimen collected from
Cape Le Grand National Park Cape Le Grand National Park is a national park in Western Australia, south-east of Perth and east of Esperance. The park covers an area of The area is an ancient landscape which has been above sea level for well over 200 million years and ...
,
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to ...
, in 2009. The name of the genus is derived from the Latin 'zephyrus', meaning 'west wind', and refers to the windy coastal habitats that species of the genus inhabit. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
is in honour of Mark Wojcieszek, who helped collect the initial specimens of this species.


Description

Like other species in its genus, ''Zephyrarchaea marki'' is a small species of spider, with a total length of in adult males. The cephalothorax is dark reddish-brown, while the abdomen is mottled grey-brown and beige, with reddish-brown dorsal scute and sclerites. The legs are tan brown with darker ring-like markings. The "neck" is wide, with the highest point of the pars cephalica (portion of the cephalothorax between the front radial furrows) approaching the posterior third of the head. The carapace is short, measuring long, tall, and wide. It has a pronounced concave depression anterior to the highest point of the pars cephalica. The "head" is not strongly elevated dorsally; the post-ocular ratio, the length of the "head" posterior to the anterior median eyes (AME) relative to the dorsal elevation of the pars cephalica above the level of the AME, is 0.21. The chelicerae have proximal tufts and an additional comb of accessory setae on front face of the paturon. The abdomen is long and wide, and almost spherical when viewed from the side. It lacks hump-like tubercles along its back but has highly flattened mound-like vestiges. The dorsal scutes are fused anteriorly to the epigastric sclerites, extending posteriorly to cover nearly anterior two-thirds of dorsal abdomen. Pedipalps are pyriform (pear-shaped) when unexpanded, with a broad, distally curved embolus supported by conductor sclerites 1 and 2. Tegular sclerite 1 extends horizontally and is strongly curved in when viewed prolaterally, with a flattened, broadly rounded, paddle-shaped apex. Tegular sclerites 2 and 3 project beyond the retro-distal rim of the tegulum. The femur of leg 1 is long; the ratio of this femur and the carapace is 1.95. The appearance of the female has not been described. ''Zephyrarchaea'' spiders have relatively uniform and cryptic coloration, generally only showing subtle intraspecific variation in abdominal patterning. ''Z. marki'' can be distinguished from ''Z. janineae'' and ''Z. mainae'' by the absence of dorsal hump-like tubercles on the abdomen, and from ''Z. marae'' and ''Z. vichickmani'' by the presence of a proximal tuft of accessory setae on the male chelicerae. ''Z. barrettae'' and ''Z. melindae'' differ in the shape of tegular sclerites 2 and 3; these project well beyond the retro-distal rim of the tegulum in ''Z. marki''. ''Z. porchi'' has a smaller, less protuberant proximal bulge on the male chelicerae.


Distribution and conservation

The species is known only from its
type locality Type locality may refer to: * Type locality (biology) * Type locality (geology) See also * Local (disambiguation) * Locality (disambiguation) {{disambiguation ...
of Thistle Cove at Cape Le Grand National Park. Collected specimens were found in elevated leaf litter in a dense coastal thickets of ''
Banksia speciosa ''Banksia speciosa'', commonly known as the showy banksia, is a species of large shrub or small tree in the family Proteaceae. It is found on the south coast of Western Australia between Hopetoun (33°57′ S) and the Great Australian Bi ...
'', behind a beach. ''Zephyrarchaea marki'' has not yet been evaluated and assigned a conservation status by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natur ...
. However, the species is endemic to a very small range and its only known population may be threatened by fire, dieback disease affecting ''Banksia'', and climate change.


References

{{Taxonbar, from=Q18667419 Spiders described in 2012 Archaeidae