Zemaciinae
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Borsoniidae is a monophyletic
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of small to medium-sized sea snails, marine gastropod
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s in the superfamily
Conoidea Conoidea is a superfamily of predatory sea snails, marine gastropod mollusks within the suborder Hypsogastropoda. This superfamily is a very large group of marine mollusks, estimated at about 340 recent valid genera and subgenera, and conside ...
.Bouchet, P. (2011). Borsoniidae. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=153870 on 2011-08-12Bouchet P. & Rocroi J.-P. (Ed.) (2005). "Classification and nomenclator of gastropod families". ''
Malacologia ''Malacologia'' is a peer-reviewed scientific journal in the field of malacology, the study of mollusks. The journal publishes articles in the fields of molluscan systematics, ecology, population ecology, genetics, molecular genetics, evolu ...
'' 47(1-2). . 397 pp.
In 2011, Bouchet, Kantor ''et al''. brought genera from the subfamilies Clathurellinae and
Raphitominae Raphitominae was a subfamily of small to quite large sea snails, marine gastropod mollusks in the superfamily Conoidea. Bouchet, Kantor ''et al''. elevated in 2011 the subfamily Raphitominae (which at that point had been placed in the family ...
they were previously placed in the family Conidae), as well as genera from the subfamily Zemaciinae (at that point belonging to the family
Turridae Turridae is a taxonomic family name for a number of predatory sea snails, marine gastropod mollusks in the superfamily Conoidea. MolluscaBase (2018). Turridae H. Adams & A. Adams, 1853 (1838). Accessed through: World Register of Marine Specie ...
), together to form the family Borsoniidae. This re-arrangement was based on anatomical characters and a dataset of molecular sequences of three
gene In biology, the word gene (from , ; "... Wilhelm Johannsen coined the word gene to describe the Mendelian units of heredity..." meaning ''generation'' or ''birth'' or ''gender'') can have several different meanings. The Mendelian gene is a b ...
fragments Bouchet, P.; Kantor, Y. I.; Sysoev, A.; Puillandre, N. (2011). ''A new operational classification of the Conoidea''. Journal of Molluscan Studies. 77, 273-308
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Description

This family is a rather heterogenous group, with wide-ranging varieties in their properties. The medium- to large-sized shells are fusiform to biconic in shape. Their size has a wide range (between 5 mm and 80 mm). The shell can be longitudinally coarsely ribbed, but axial ribs are sometimes obsolete to absent. The smooth
columella Lucius Junius Moderatus Columella (; Arabic: , 4 – ) was a prominent writer on agriculture in the Roman Empire. His ' in twelve volumes has been completely preserved and forms an important source on Roman agriculture, together with the wo ...
has a strong (one to three plicae) to obsolete plication upon the middle. This plication varies within genera. In the genus ''
Cordiera ''Cordiera'' is a genus of flowering plants in the family Rubiaceae. The genus is found in central and southern tropical America. Species * '' Cordiera concolor'' ( Cham.) Kuntze ** ''Cordiera concolor'' var. ''concolor'' ** ''Cordiera concolo ...
'' there are two columellar plaits. The
aperture In optics, an aperture is a hole or an opening through which light travels. More specifically, the aperture and focal length of an optical system determine the cone angle of a bundle of rays that come to a focus in the image plane. An ...
is elliptical to oval in shape and it has a short to moderately long (e.g. '' Zemacies excelsa'') siphonal canal and a deep anal sinus. The anterior canal has a moderate length and is slightly twisted to the left. The thin outer lip is arcuate. The vestigial or fully developed operculum has a terminal nucleus, and may be missing in some species. The radula is absent in the genus '' Zemacies''. McLean, J.H. (1971) ''A revised classification of the family Turridae, with the proposal of new subfamilies, genera, and subgenera from the Eastern Pacific''. The Veliger, 14, 114–130
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Genera

* '' Antarctospira'' Kantor, Harasewych & Puillandre, 2016 * ? '' Apaturris'' Iredale, 1917 * '' Aphanitoma'' Bellardi, 1875 * '' Asthenotoma'' Harris & Burrows, 1891 * '' Austroturris'' Laseron, 1954 * '' Awateria'' Suter, 1917 * '' Bathytoma'' Harris & Burrows, 1891 * '' Belaturricula'' Powell, 1951 * ''
Borsonella ''Borsonella'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae. Description (Original description) The shell has a small, blunt nucleus of one or two whorls. The sculpture is chiefly spiral, feeble, except for one ...
'' Dall, 1908 * '' Borsonellopsis'' McLean, 1971 * '' Borsonia'' Bellardi, 1839 * '' Cordieria'' Rouault, 1848 * '' Darbya'' Bartsch, 1934 * '' Diptychophlia'' Berry, 1964 * ''
Drilliola ''Drilliola'' is a genus of sea snails, marine gastropod mollusks in the family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The p ...
'' Locard, 1897 * '' Filodrillia'' Hedley, 1922 * '' Genota'' H. Adams & A. Adams, 1853 * ''
Glyptaesopus ''Glyptaesopus'' is a genus of sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Borsoniidae Description The small shell is slender and contains 5½ sculptured Whorl (mollusc), whorls with weakly incised Sutur ...
'' Pilsbry & Olsson, 1941 * '' Heteroturris'' Powell, 1967 * '' Maoritomella'' Powell, 1942 * '' Microdrillia'' Casey, 1903 * '' Ophiodermella'' Bartsch, 1944 * '' Paraborsonia'' Pilsbry, 1922 * '' Phenatoma'' Finlay, 1924 * '' Pulsarella'' Laseron, 1954 * '' Retidrillia'' McLean, 2000 * ''
Suavodrillia ''Suavodrillia'' is a genus of sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Borsoniidae. Species Species within the genus ''Suavodrillia'' include: * ''Suavodrillia declivis'' (Martens, 1880) * ''Suavodri ...
'' Dall, 1918 * '' Tomopleura'' Casey, 1904 * '' Tropidoturris'' Kilburn, 1986 * ''
Typhlodaphne ''Typhlodaphne'' is a genus of sea snails, marine gastropod mollusks in the family (biology), family Borsoniidae. Description The smooth protoconch is bluntly rounded and paucispiral, consisting of two Whorl (mollusc), whorls. The sinus is subsu ...
'' Powell, 1951 * ''
Typhlomangelia ''Typhlomangelia'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae, the cone snails A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, ...
'' G.O. Sars, 1878 * '' Zemacies'' Finlay, 1926 ;Genera brought into synonymy : * ''Acrobela'' Thiele, 1925 : synonym of '' Microdrillia'' Casey, 1903 * ''Acropota'' F. Nordsieck, 1977: synonym of '' Microdrillia'' Casey, 1903 * ''Boettgeria'' Peyrot, 1931 : synonym of ''Boettgeriola'' Wenz, 1943 * ''Boettgeriola'' Wenz, 1943 : synonym of '' Borsonia'' Bellardi, 1839 * ''Genotia'' P. Fischer, 1883 : synonym of '' Genota'' H. Adams & A. Adams, 1853 * ''Micantapex'' Iredale, 1936 : synonym of '' Bathytoma'' Harris & Burrows, 1891 * ''Narraweena (gastropod)'' Laseron, 1954 : synonym of '' Maoritomella'' Powell, 1942 * ''Oligotoma'' Bellardi, 1875 : synonym of '' Asthenotoma'' Harris & Burrows, 1891 * ''Parabathytoma'' Shuto, 1961 : synonym of '' Bathytoma'' Harris & Burrows, 1891 * ''Riuguhdrillia'' Oyama, 1951 : synonym of '' Bathytoma'' Harris & Burrows, 1891 * ''Vexithara'' Finlay, 1926 : synonym of ''
Typhlomangelia ''Typhlomangelia'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae, the cone snails A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, ...
'' G.O. Sars, 1878 * ''Viridoturris'' Powell, 1964 : synonym of ''
Typhlomangelia ''Typhlomangelia'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae, the cone snails A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, ...
'' G.O. Sars, 1878


References


External links


Gastropods.com: Borsoniidae; accessed : 12 August 2011
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{{Taxonbar, from=Q4946684 Conoidea