Zelkova × Verschaffeltii
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''Zelkova'' × ''verschaffeltii'' (Dippel) G.Nicholson (cut-leaf zelkova) is a zelkova
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
of
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
origin. It was originally described in 1892 by Leopold Dippel from a cultivated plant as ''Zelkova japonica'' var. ''verschaffeltii'', suggesting an eastern Asian origin. In 1896, George Nicholson raised it to species rank as ''Zelkova verschaffeltii'', while its much closer similarity to ''
Zelkova carpinifolia ''Zelkova carpinifolia'', known as Caucasian zelkova, Caucasian elm or just zelkova, is a species of ''Zelkova'', native to the Caucasus, Kaçkar, and Alborz mountains in the extreme southeast of Europe and southwest Asia. It is a medium-size ...
'' led
Augustine Henry Augustine Henry (2 July 1857 – 23 March 1930) was a British-born Irish plantsman and sinologist. He is best known for sending over 15,000 dry specimens and seeds and 500 plant samples to Kew Gardens in the United Kingdom. By 1930, he was a rec ...
to suggest it might be a hybrid between ''Zelkova carpinifolia'' and ''
Zelkova abelicea ''Zelkova abelicea'' is a species of tree in the family Ulmaceae. It is referred to by the common names Cretan zelkova, and on Crete proper as abelitsia (αμπελιτσιά). It is endemic to Crete. It is found in small numbers and is classifie ...
''.Bean, W. J. (1980). ''Trees and Shrubs Hardy in the British Isles'' 8th ed., vol. 4. John Murray . More recent authors most widely regarded it as a cultivar, either not ascribed to any ''Zelkova'' species in particular, or placed under ''Z. carpinifolia'' with no suggestion of hybrid origin.Czerepanov, S. (1957). Revisio specierum generum Zelkova Spach et Hemiptelea Planchon. ''Bot. Material'' 18: 58-72. Analysis of
flavonoid Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...
s has subsequently suggested that it is a hybrid between ''Zelkova carpinifolia'' and '' Zelkova serrata'' : "The chromatographic evidence would suggest that ''Z. X verschaffeltii'' is a hybrid between ''Z. carpinifolia'' and ''Z. serrata''. ''Z. serrata'' was first introduced into England in 1861, and ''Z. verschaffeltii'' has been cultivated since 1886. It may well have been that some of the ''Z. serrata'' trees reached sexual maturity before 1886 and crossed with ''Z. carpinifolia'' already in cultivation. The fact that ''Z. X verschaffeltii'' is really a clone would indicate that such a cross might be a rare event, and our experience would tend to substantiate this hypothesis."Santamour, F. S. (1983). Flavonoid distribution in ''Zelkova''. ''J. Arboriculture'' 9 (7): 190-192.


Name in error

The tree was originally misidentified as an elm. The putative
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
''Ulmus'' 'Pitteurs Pendula' was listed by C. de Vos in ''Handboek'' Supplement 16, 1890, as ''Ulmus pitteursii pendula''. However,
Hans M. Heybroek Hans M. Heybroek (born 1927, Bussum, died 2022, Wassenaar)Interview with Hans Heybro ...
, erstwhile head of the Dutch elm breeding programme at the de Dorschkamp Research Institute for Forestry and Landscape Planning at
Wageningen Wageningen () is a municipality and a historic city in the central Netherlands, in the province of Gelderland. It is famous for Wageningen University, which specialises in life sciences. The municipality had a population of in , of which many t ...
, identified the tree as ''Zelkova × verschaffeltii''.


Description

A large
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
or small, bushy-crowned
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
< 15 m tall, with mottled grey and brown
bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, ...
and graceful habit. The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are narrow, 3.5–8 cm long and 1.3–6.6 cm broad, deeply incised with between five and ten pairs of triangular teeth. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
is rarely produced, a small two-lobed dry nut-like
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
4–5 mm diameter.Andrews, S. (1994). Tree of the year: Zelkova. ''Int. Dendrol. Soc. Yearbook'' 1993: 11-30.Mitchell, A. F. (1974). ''A Field Guide to the Trees of Britain and Northern Europe''. Collins Mitchell, A. F. (1982). ''The Trees of Britain and Northern Europe''. Collins White, J. & More, D. (2003). ''Trees of Britain & Northern Europe''. Cassell's, London.


Cultivation

First found in cultivation in the
van Houtte Van Houtte Inc. is a company based in Montreal, Quebec, Canada that processes, distributes and sells coffee and coffee related goods. They have expanded their product selection, which now includes tropical teas, sandwiches and salads. It was founde ...
nursery at
Ghent Ghent ( nl, Gent ; french: Gand ; traditional English: Gaunt) is a city and a municipality in the Flemish Region of Belgium. It is the capital and largest city of the East Flanders province, and the third largest in the country, exceeded in ...
in
Belgium Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to th ...
before 1885.


Etymology

The tree is named for the Belgian botanist
Ambroise Verschaffelt Ambroise Colette Alexandre Verschaffelt (11 December 1825 – 16 May 1886) was a distinguished Belgian horticulturist and author. His grandfather Pierre-Antoine (1764–1844) was amongst the founders of the "Floralies gantoises" in 1808. His f ...
who owned the Verschaffelt Nursery in
Ghent Ghent ( nl, Gent ; french: Gand ; traditional English: Gaunt) is a city and a municipality in the Flemish Region of Belgium. It is the capital and largest city of the East Flanders province, and the third largest in the country, exceeded in ...
, later acquired by Jean Jules Linden, the renowned horticulturist and
camellia ''Camellia'' (pronounced or ) is a genus of flowering plants in the family Theaceae. They are found in eastern and southern Asia, from the Himalayas east to Japan and Indonesia. There are more than 220 described species, with some controversy ...
specialist.


Accessions


Europe

* Royal Botanic Garden Edinburgh, acc. no. 19531019 (15 m tall in 2004Tree Register of the British Isles) *
Sir Harold Hillier Gardens The Sir Harold Hillier Gardens is an arboretum comprising 72 hectares (180 acres) accommodating over 42,000 trees and shrubs in about 12,000 taxa, notably a collection of oaks, camellia, magnolia and rhododendron. The Gardens are located nor ...
, acc. no. 1982.0194, as ''Z. carpinifolia'' 'Verschaffeltii'. *
University of Copenhagen The University of Copenhagen ( da, Københavns Universitet, KU) is a prestigious public university, public research university in Copenhagen, Copenhagen, Denmark. Founded in 1479, the University of Copenhagen is the second-oldest university in ...
Botanic Garden, acc. no. P.1977-5020, as ''Z. carpinifolia'' 'Verschaffeltii'. *
Icomb Place Icomb Place (pronounced "Ickum") is a medieval manor house on the edge of the village of Icomb, near Stow on the Wold in Gloucestershire. The word "place" in this context is thought to be a precursor of "palace". Description The house is a Grade ...
gardens, as ''Z. carpinifolia'' 'Verschaffeltii'. * Wijdemeren city counsel, The Netherlands, Kuijerpark, Nederhorst den Berg as Z x verschaffeltii


North America

*
Longwood Gardens Longwood Gardens is a botanical garden that consists of over 1,077 acres (436 hectares; 4.36 km2) of gardens, woodlands, and meadows in Kennett Square, Pennsylvania, United States in the Brandywine Creek Valley. It is one of the premier h ...
, acc. no. 1992-0138, as ''Z. carpinifolia'' 'Verschaffeltii'. * New York Botanical Garden, acc. no. 2793/93, as ''Z. verschaffeltii''.


Synonymy

* ''Ulmus'' 'Pitteurs Pendula', in error.


References

{{DEFAULTSORT:Zelkova verschaffeltii verschaffeltii Misidentified elm cultivar Hybrid plants