Zebras (, ) (subgenus ''Hippotigris'') are African
equines
''Equus'' , is a genus of mammals in the family (biology), family Equidae, which includes horses, donkeys, and zebras. Within the Equidae, ''Equus'' is the only recognized Extant taxon, extant genus, comprising seven living species. Like Equida ...
with distinctive black-and-white striped
coat
A coat typically is an outer garment for the upper body as worn by either gender for warmth or fashion. Coats typically have long sleeves and are open down the front and closing by means of buttons, zippers, hook-and-loop fasteners, toggles, a ...
s. There are three
living
Living or The Living may refer to:
Common meanings
*Life, a condition that distinguishes organisms from inorganic objects and dead organisms
** Living species, one that is not extinct
*Personal life, the course of an individual human's life
* ...
species: the
Grévy's zebra (''Equus grevyi''),
plains zebra
The plains zebra (''Equus quagga'', formerly ''Equus burchellii''), also known as the common zebra, is the most common and geographically widespread species of zebra. Its range is fragmented, but spans much of southern and eastern Africa south of ...
(''E. quagga''), and the
mountain zebra
The mountain zebra (''Equus zebra'') is a zebra species in the family Equidae, native to southwestern Africa. There are two subspecies, the Cape mountain zebra (''E. z. zebra'') found in South Africa and Hartmann's mountain zebra (''E. z. hartman ...
(''E. zebra''). Zebras share the genus ''Equus'' with
horses
The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
and
asses
Ass most commonly refers to:
* Buttocks (in informal American English)
* Donkey or ass, ''Equus africanus asinus''
**any other member of the subgenus ''Asinus''
Ass or ASS may also refer to:
Art and entertainment
* Ass (album), ''Ass'' (album ...
, the three groups being the only living members of the family
Equidae
Equidae (sometimes known as the horse family) is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. All extant species are in the genus '' Equus'', ...
. Zebra stripes come in different patterns, unique to each individual. Several theories have been proposed for the function of these stripes, with most evidence supporting them as a deterrent for biting flies. Zebras inhabit
eastern
Eastern may refer to:
Transportation
*China Eastern Airlines, a current Chinese airline based in Shanghai
*Eastern Air, former name of Zambia Skyways
*Eastern Air Lines, a defunct American airline that operated from 1926 to 1991
*Eastern Air Li ...
and
southern Africa
Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number of ...
and can be found in a variety of habitats such as
savannahs,
grasslands
A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natural ...
, woodlands,
shrubland
Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It m ...
s, and mountainous areas.
Zebras are primarily
grazers and can subsist on lower-quality vegetation. They are preyed on mainly by
lions
The lion (''Panthera leo'') is a large cat of the genus ''Panthera'' native to Africa and India. It has a muscular, broad-chested body; short, rounded head; round ears; and a hairy tuft at the end of its tail. It is sexually dimorphic; adult ...
, and typically flee when threatened but also bite and kick. Zebra species differ in
social behaviour
Social behavior is behavior among two or more organisms within the same species, and encompasses any behavior in which one member affects the other. This is due to an interaction among those members. Social behavior can be seen as similar to an ...
, with plains and mountain zebra living in stable
harems
Harem (Persian: حرمسرا ''haramsarā'', ar, حَرِيمٌ ''ḥarīm'', "a sacred inviolable place; harem; female members of the family") refers to domestic spaces that are reserved for the women of the house in a Muslim family. A hare ...
consisting of an adult male or
stallion
A stallion is a male horse that has not been gelded (castrated).
Stallions follow the conformation and phenotype of their breed, but within that standard, the presence of hormones such as testosterone may give stallions a thicker, "cresty" nec ...
, several adult females or
mares
A mare is an adult female horse or other equine. In most cases, a mare is a female horse over the age of three, and a filly is a female horse three and younger. In Thoroughbred horse racing, a mare is defined as a female horse more than four y ...
, and their young or
foals
A foal is an equine up to one year old; this term is used mainly for horses, but can be used for donkeys. More specific terms are colt for a male foal and filly for a female foal, and are used until the horse is three or four. When the foal is ...
; while Grévy's zebra live alone or in loosely associated herds. In harem-holding species, adult females mate only with their harem stallion, while male Grévy's zebras establish
territories
A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal.
In international politics, a territory is usually either the total area from which a state may extract power resources or an ...
which attract females and the species is
promiscuous
Promiscuity is the practice of engaging in sexual activity frequently with different Sexual partner, partners or being indiscriminate in the choice of sexual partners. The term can carry a moral judgment. A common example of behavior viewed as pro ...
. Zebras communicate with various vocalisations, body postures and facial expressions.
Social grooming
Social grooming is a behavior in which social animals, including humans, clean or maintain one another's body or appearance. A related term, allogrooming, indicates social grooming between members of the same species. Grooming is a major soci ...
strengthens social bonds in plains and mountain zebras.
Zebras' dazzling stripes make them among the most recognisable mammals. They have been featured in art and stories in Africa and beyond. Historically, they have been highly sought after by exotic animal collectors, but unlike
horses
The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
and
donkeys
The domestic donkey is a hoofed mammal in the family Equidae, the same family as the horse. It derives from the African wild ass, ''Equus africanus'', and may be classified either as a subspecies thereof, ''Equus africanus asinus'', or as a ...
, zebras have never been truly
domesticated
Domestication is a sustained multi-generational relationship in which humans assume a significant degree of control over the reproduction and care of another group of organisms to secure a more predictable supply of resources from that group. A ...
. The
International Union for Conservation of Nature
The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
(IUCN) lists the Grévy's zebra as
endangered
An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inva ...
, the mountain zebra as
vulnerable and the plains zebra as
near-threatened
A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify fo ...
. The
quagga
The quagga ( or ) (''Equus quagga quagga'') is a subspecies of the plains zebra that was endemic to South Africa until it was hunted to extinction in the late 19th century. It was long thought to be a distinct species, but early genetic ...
, a type of plains zebra, was driven to extinction in the 19th century. Nevertheless, zebras can be found in numerous protected areas.
Etymology
The English name "zebra" derives from
Italian
Italian(s) may refer to:
* Anything of, from, or related to the people of Italy over the centuries
** Italians, an ethnic group or simply a citizen of the Italian Republic or Italian Kingdom
** Italian language, a Romance language
*** Regional Ita ...
,
Spanish
Spanish might refer to:
* Items from or related to Spain:
**Spaniards are a nation and ethnic group indigenous to Spain
**Spanish language, spoken in Spain and many Latin American countries
**Spanish cuisine
Other places
* Spanish, Ontario, Cana ...
or
Portuguese
Portuguese may refer to:
* anything of, from, or related to the country and nation of Portugal
** Portuguese cuisine, traditional foods
** Portuguese language, a Romance language
*** Portuguese dialects, variants of the Portuguese language
** Portu ...
.
Its origins may lie in the
Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''equiferus'', meaning "wild horse". ''Equiferus'' appears to have entered into Portuguese as ''ezebro'' or ''zebro'', which was originally used for a legendary equine in the wilds of the
Iberian Peninsula
The Iberian Peninsula (),
**
* Aragonese and Occitan: ''Peninsula Iberica''
**
**
* french: Péninsule Ibérique
* mwl, Península Eibérica
* eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
during the Middle Ages. In 1591, Italian explorer
Filippo Pigafetta Filippo Pigafetta (1533–1604) was an Italian mathematician and explorer.
Pigafetta's ''Relatione del reame del Congo'' (A Report of the Kingdom of Congo and of the Surrounding Countries) 1591 was translated into English, Latin (as ''Regnum Congo ...
recorded "zebra" being used to refer to the African animals by Portuguese visitors to the continent.
In ancient times, the African zebra was called ''hippotigris'' ("horse tiger") by the
Greeks
The Greeks or Hellenes (; el, Έλληνες, ''Éllines'' ) are an ethnic group and nation indigenous to the Eastern Mediterranean and the Black Sea regions, namely Greece, Cyprus, Albania, Italy, Turkey, Egypt, and, to a lesser extent, oth ...
and
Romans
Roman or Romans most often refers to:
*Rome, the capital city of Italy
* Ancient Rome, Roman civilization from 8th century BC to 5th century AD
*Roman people, the people of ancient Rome
*''Epistle to the Romans'', shortened to ''Romans'', a lette ...
.
The word "zebra" was traditionally pronounced with a long initial vowel, but over the course of the 20th century the pronunciation with the short initial vowel became the norm in the UK and the
Commonwealth
A commonwealth is a traditional English term for a political community founded for the common good. Historically, it has been synonymous with "republic". The noun "commonwealth", meaning "public welfare, general good or advantage", dates from the ...
. The pronunciation with a long initial vowel remains standard in
US English
American English, sometimes called United States English or U.S. English, is the set of varieties of the English language native to the United States. English is the most widely spoken language in the United States and in most circumstances i ...
.
Taxonomy
Zebras are classified in the genus ''
Equus'' (known as equines) along with
horses
The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
and
asses
Ass most commonly refers to:
* Buttocks (in informal American English)
* Donkey or ass, ''Equus africanus asinus''
**any other member of the subgenus ''Asinus''
Ass or ASS may also refer to:
Art and entertainment
* Ass (album), ''Ass'' (album ...
. These three groups are the only living members of the family
Equidae
Equidae (sometimes known as the horse family) is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. All extant species are in the genus '' Equus'', ...
.
The
plains zebra
The plains zebra (''Equus quagga'', formerly ''Equus burchellii''), also known as the common zebra, is the most common and geographically widespread species of zebra. Its range is fragmented, but spans much of southern and eastern Africa south of ...
and
mountain zebra
The mountain zebra (''Equus zebra'') is a zebra species in the family Equidae, native to southwestern Africa. There are two subspecies, the Cape mountain zebra (''E. z. zebra'') found in South Africa and Hartmann's mountain zebra (''E. z. hartman ...
were traditionally placed in the
subgenus
In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus.
In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
''Hippotigris'' (C. H. Smith, 1841) in contrast to the
Grévy's zebra which was considered the sole species of subgenus ''Dolichohippus'' (Heller, 1912).
Groves and Bell (2004) placed all three species in the subgenus ''Hippotigris''.
A 2013
phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
study found that the plains zebra is more closely related to Grévy's zebras than mountain zebras.
The extinct
quagga
The quagga ( or ) (''Equus quagga quagga'') is a subspecies of the plains zebra that was endemic to South Africa until it was hunted to extinction in the late 19th century. It was long thought to be a distinct species, but early genetic ...
was originally classified as a distinct species. Later genetic studies have placed it as the same species as the plains zebra, either a subspecies or just the southernmost population.
Molecular evidence supports zebras as a
monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
lineage
Lineage may refer to:
Science
* Lineage (anthropology), a group that can demonstrate its common descent from an apical ancestor or a direct line of descent from an ancestor
* Lineage (evolution), a temporal sequence of individuals, populati ...
.
''Equus'' originated in North America and direct
paleogenomic sequencing of a 700,000-year-old middle Pleistocene horse
metapodial Metapodials are long bones of the hand (metacarpals) and feet (metatarsal
The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. ...
bone from Canada implies a date of 4.07 million years ago (mya) for the most
recent common ancestor
In biology and genetic genealogy, the most recent common ancestor (MRCA), also known as the last common ancestor (LCA) or concestor, of a set of organisms is the most recent individual from which all the organisms of the set are descended. The ...
of the equines within the range of 4.0 to 4.5 mya.
Horses
split
Split(s) or The Split may refer to:
Places
* Split, Croatia, the largest coastal city in Croatia
* Split Island, Canada, an island in the Hudson Bay
* Split Island, Falkland Islands
* Split Island, Fiji, better known as Hạfliua
Arts, enterta ...
from asses and zebras around this time and equines colonised Eurasia and Africa around 2.1–3.4 mya. Zebras and asses diverged from each other close to 2 mya. The mountain zebra diverged from the other species around 1.6 mya and the plains and Grévy's zebra split 1.4 mya.
![Equus quagga quagga, coloured](https://upload.wikimedia.org/wikipedia/commons/c/cd/Equus_quagga_quagga%2C_coloured.jpg)
The cladogram of ''Equus'' below is based on Vilstrup and colleagues (2013) and Jónsson and colleagues (2014):
[
]
Living species
Fossil record
In addition to the three living species, some fossil zebras and relatives have also been identified. ''Equus koobiforensis'' is an early equine basal to zebras found in the Shungura Formation
The Shungura Formation is a stratigraphic formation located in the Omo river basin in Ethiopia. It dates to the Late Pliocene to Early Pleistocene. Oldowan tools have been found in the formation, suggesting early use of stone tools by hominins. A ...
, Ethiopia
Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
and the Olduvai Gorge
The Olduvai Gorge or Oldupai Gorge in Tanzania is one of the most important paleoanthropology, paleoanthropological localities in the world; the many sites exposed by the gorge have proven invaluable in furthering understanding of early human ev ...
, Tanzania
Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
, and dated to around 2.3 mya. ''E. oldowayensis'' is identified from remains in Olduvai Gorge dating to 1.8 mya. Fossil skulls of ''E. mauritanicus'' from Algeria which date to around 1 mya appears to show affinities with the plains zebra. '' E. capensis'', known as the Cape zebra, appeared around 2 mya and lived throughout southern and eastern Africa.
Non-African equines that may have been basal to zebras include ''E. sansaniensis'' of Eurasia (circa 2.5 mya) and '' E. namadicus'' (circa 2.5 mya) and '' E. sivalensis'' (circa 2.0 mya) of the Indian subcontinent. A 2017 mitochondrial DNA study placed the Eurasian ''E. ovodovi'' and the subgenus ''Sussemionus'' lineage as closer to zebras than to asses.
Hybridisation
Fertile hybrids have been reported in the wild between plains and Grévy's zebra. Hybridisation has also been recorded between the plains and mountain zebra, though it is possible that these are infertile due to the difference in chromosome numbers between the two species. Captive zebras have been bred with horses and donkeys
The domestic donkey is a hoofed mammal in the family Equidae, the same family as the horse. It derives from the African wild ass, ''Equus africanus'', and may be classified either as a subspecies thereof, ''Equus africanus asinus'', or as a ...
; these are known as zebroids. A zorse is a cross between a zebra and a horse; a zonkey between a zebra and a donkey and a zoni between a zebra and a pony
A pony is a type of small horse ('' Equus ferus caballus''). Depending on the context, a pony may be a horse that is under an approximate or exact height at the withers, or a small horse with a specific conformation and temperament. Compared ...
. Zebroids are often born sterile with dwarfism
Dwarfism is a condition wherein an organism is exceptionally small, and mostly occurs in the animal kingdom. In humans, it is sometimes defined as an adult height of less than , regardless of sex; the average adult height among people with dw ...
.
Characteristics
As with all wild equines, zebra have barrel-chested bodies with tufted tails, elongated faces and long necks with long, erect manes
In ancient Roman religion, the ''Manes'' (, , ) or ''Di Manes'' are chthonic deities sometimes thought to represent souls of deceased loved ones. They were associated with the ''Lares'', ''Lemures,'' '' Genii'', and ''Di Penates'' as deities ('' ...
. Their thin legs are each supported by a spade-shaped toe covered in a hard hoof
The hoof (plural: hooves) is the tip of a toe of an ungulate mammal, which is covered and strengthened with a thick and horny keratin covering. Artiodactyls are even-toed ungulates, species whose feet have an even number of digits, yet the rumin ...
. Their dentition
Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolo ...
is adapted for grazing
In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to roam around and consume wild vegetations in order to convert the otherwise indigestible (by human gut) cellulose within grass and other ...
; they have large incisors that clip grass blades and rough molars and premolars well suited for grinding. Males have spade-shaped canines, which can be used as weapons in fighting. The eyes of zebras are at the sides and far up the head, which allows them to look over the tall grass while feeding. Their moderately long, erect ears are movable and can locate the source of a sound.
Unlike horses, zebras and asses have chestnut callosities present only on their front legs. In contrast to other living equines, zebra have longer front legs than back legs. Diagnostic traits of the zebra skull include: its relatively small size with a straight profile, more projected eye sockets, narrower rostrum, reduced postorbital bar The postorbital bar (or postorbital bone) is a bony arched structure that connects the frontal bone of the skull to the zygomatic arch, which runs laterally around the eye socket. It is a trait that only occurs in mammalian taxa, such as most strep ...
, separation of the metaconid
Many different terms have been proposed for features of the tooth crown in mammals.
The structures within the molars receive different names according to their position and morphology. This nomenclature was developed by Henry Fairfield Osborn ...
and metastylid of the tooth by a V-shaped canal and rounded enamel wall.
Stripes
Zebras are easily recognised by their bold black-and-white striping patterns. The coat
A coat typically is an outer garment for the upper body as worn by either gender for warmth or fashion. Coats typically have long sleeves and are open down the front and closing by means of buttons, zippers, hook-and-loop fasteners, toggles, a ...
appears to be white with black stripes, as indicated by the belly and legs when unstriped, but the skin underneath is black. Young or foal
A foal is an equine up to one year old; this term is used mainly for horses, but can be used for donkeys. More specific terms are colt for a male foal and filly for a female foal, and are used until the horse is three or four. When the foal i ...
s are born with brown and white coats, and the brown darkens with age. A dorsal
Dorsal (from Latin ''dorsum'' ‘back’) may refer to:
* Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism
* Dorsal, positioned on top of an aircraft's fuselage
* Dorsal co ...
line acts as the backbone for vertical stripes along the sides, from the head to the rump. On the snout they curve toward the nostrils, while the stripes above the front legs split into two branches. On the rump, they develop into species-specific patterns. The stripes on the legs, ears and tail are separate and horizontal.
Striping patterns are unique to an individual and heritable. During embryonic development
An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sperm ...
, the stripes appear at eight months, but the patterns may be determined at three to five weeks. For each species there is a point in embryonic development where the stripes are perpendicular to the dorsal line and spaced apart. However, this happens at three weeks of development for the plains zebra, four weeks for the mountain zebra, and five for Grévy's zebra. The difference in timing is thought to be responsible for the differences in the striping patterns of the different species.
Various abnormalities of the patterns have been documented in plains zebras. In melanistic
The term melanism refers to black pigment and is derived from the gr, μελανός. Melanism is the increased development of the dark-colored pigment melanin in the skin or hair.
Pseudomelanism, also called abundism, is another variant of pi ...
zebras, dark stripes are highly concentrated on the torso but the legs are whiter. "Spotted" individuals display disruptions in black striping patterns between the shoulders and hips. There have even been morphs with white spots on dark backgrounds. Striping abnormalities have been linked to inbreeding
Inbreeding is the production of offspring from the mating or breeding of individuals or organisms that are closely related genetically. By analogy, the term is used in human reproduction, but more commonly refers to the genetic disorders and o ...
.[ ]Albino
Albinism is the congenital absence of melanin in an animal or plant resulting in white hair, feathers, scales and skin and pink or blue eyes. Individuals with the condition are referred to as albino.
Varied use and interpretation of the term ...
zebras have been recorded in the forests of Mount Kenya
Mount Kenya (Kikuyu: ''Kĩrĩnyaga'', Kamba, ''Ki Nyaa'') is the highest mountain in Kenya and the second-highest in Africa, after Kilimanjaro. The highest peaks of the mountain are Batian (), Nelion () and Point Lenana (). Mount Kenya is locat ...
, with the dark stripes being blonde. The quagga had brown and white stripes on the head and neck, brown upper parts and a white belly, tail and legs.
Function
The function of stripes in zebras has been discussed among biologists since at least the 19th century. Popular hypotheses include the following:
* The crypsis
In ecology, crypsis is the ability of an animal or a plant to avoid observation or detection by other animals. It may be a predation strategy or an antipredator adaptation. Methods include camouflage, nocturnality, subterranean lifestyle and ...
hypothesis was proposed by Alfred Wallace
Alfred Russel Wallace (8 January 1823 – 7 November 1913) was a British naturalist, explorer, geographer, anthropologist, biologist and illustrator. He is best known for independently conceiving the theory of evolution through natural sele ...
in 1896 and suggests that the stripes allow the animal to blend in with its environment or break out its outline so predators can not perceive it as a single entity. In 1871, Charles Darwin
Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended fr ...
remarked that "the zebra is conspicuously striped, and stripes on the open plains of South Africa cannot afford any protection". Zebras graze in open habitat and do not behave cryptically, being noisy, fast, and social. They do not freeze when detecting a predator. In addition, lions and hyenas do not appear to perceive the stripes when they are certain distance away during daytime, thus making the stripes useless in disrupting the outline. Stripes also do not appear to make zebras more difficult to find than more solidly coloured animals of similar size, and predators may still be able to detect them by scent or hearing. The camouflaging stripes of woodland living ungulates like bongos
Bongos ( es, bongó) are an Afro-Cuban percussion instrument consisting of a pair of small open bottomed hand drums of different sizes. They are struck with both hands, most commonly in an eight-stroke pattern called ''martillo'' (hammer). The ...
and bushbucks are much less vivid with less contrast with the background colour. In addition, unlike tiger
The tiger (''Panthera tigris'') is the largest living cat species and a member of the genus '' Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily preys on u ...
stripes, the spatial frequencies of zebra stripes do not line up with their environment. A 2014 study could not find any correlations between striping patterns and woodland habitats.
* The confusion hypothesis states that the stripes confuse predators, be it by: making it harder to distinguish individuals in a group as well as determining the number of zebras in a group; making it difficult to determine an individual's outline when the group runs away; reducing a predator's ability to keep track of a target during a chase; dazzling an assailant so they have difficulty making contact; or making it difficult for a predator to deduce the zebra's size, speed and direction via motion dazzle
Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
. This theory has been proposed by several biologists since at least the 1970s. A 2014 computer study of zebra stripes found that they may create a wagon-wheel effect
The wagon-wheel effect (alternatively called stagecoach-wheel effect or stroboscopic effect) is an optical illusion in which a spoked wheel appears to rotate differently from its true rotation. The wheel can appear to rotate more slowly than the ...
and/or barber pole illusion
The barberpole illusion is a visual illusion that reveals biases in the processing of visual motion in the human brain. This visual illusion occurs when a diagonally striped pole is rotated around its vertical axis (horizontally), it appears as th ...
when in motion. The researchers concluded that this could be used against mammalian predators or biting flies. The use of the stripes for confusing mammalian predators has been questioned. The stripes of zebras could make groups seem smaller, and thus more likely to be attacked. Zebras also tend to scatter when fleeing from attackers and thus the stripes could not break up an individual's outline. Lions, in particular, appear to have no difficulty targeting and catching zebras when they get close and take them by ambush. In addition, no correlations have been found between the amount of stripes and populations of mammal predators.
* The aposematic
Aposematism is the advertising by an animal to potential predators that it is not worth attacking or eating. This unprofitability may consist of any defences which make the prey difficult to kill and eat, such as toxicity, venom, foul taste or ...
hypothesis suggests that the stripes serve as warning colouration. This hypothesis was first suggested by Wallace in 1867 and discussed in more detail by Edward Bagnall Poulton
Sir Edward Bagnall Poulton, FRS HFRSE FLS (27 January 1856 – 20 November 1943) was a British evolutionary biologist, a lifelong advocate of natural selection through a period in which many scientists such as Reginald Punnett doubted its i ...
in 1890. As with known aposematic mammals, zebras are recognisable up close, live in more open environments, have a high risk of predation and do not hide or act inconspicuous. However they are frequently preyed on by lions, suggesting that stripes do not work on them but may on smaller predators, and are not slow-moving enough to need to ward off threats. They also do not possess adequate defenses to back up the warning pattern.
* The social function hypothesis states that stripes serve a role in intraspecific or individual recognition, social bonding, mutual grooming or a signal of fitness. Darwin wrote in 1871 that "a female zebra would not admit the addresses of a male ass until he was painted so as to resemble a zebra" while Wallace stated in 1871 that: "The stripes therefore may be of use by enabling stragglers to distinguish their fellows at a distance". Regarding species and individual identification, zebras have limited range overlap with each other and horses can recognise each other using visual communication. In addition, no correlation has been found between striping and social behaviour or group numbers among equines, and no link has been found between fitness and striping.
* The thermoregulatory
Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperature ...
hypothesis suggests that stripes help to control a zebra's body temperature. In 1971, biologist H. A. Baldwin noted that heat would be absorbed by the black stripes and reflected by the white ones. In 1990, zoologist Desmond Morris
Desmond John Morris FLS ''hon. caus.'' (born 24 January 1928) is an English zoologist, ethologist and surrealist painter, as well as a popular author in human sociobiology. He is known for his 1967 book ''The Naked Ape'', and for his televisi ...
suggested that the stripes create convection current
Convection is single or multiphase fluid flow that occurs spontaneously due to the combined effects of material property heterogeneity and body forces on a fluid, most commonly density and gravity (see buoyancy). When the cause of the convect ...
s which bring down the animal's body temperature. A study from 2015 determined that environmental temperature is a strong predictor for zebra striping patterns. A 2019 study supported this, finding that where the faster air currents of the warmer black stripes meet those of the white, air swirls form. During the hottest times of the day, zebras may erect the black hair to release heat from the skin and flatten it to retain heat when it gets cooler. Others have found no evidence that zebras have lower body temperatures than other ungulates whose habitat they share, or that striping correlates with temperature. A 2018 experimental study which dressed water-filled metal barrels in horse, zebra and cattle hides found that zebra stripes have no effect on thermoregulation.
* The fly protection hypothesis holds that the stripes deter biting flies. Horse flies, in particular, spread diseases that are lethal to equines such as African horse sickness
African horse sickness (AHS) is a highly infectious and deadly disease caused by ''African horse sickness virus''. It commonly affects horses, mules, and donkeys. It is caused by a virus of the genus ''Orbivirus'' belonging to the family ''Reov ...
, equine influenza
Equine influenza (horse flu) is the disease caused by strains of influenza A that are enzootic in horse species. Equine influenza occurs globally, previously caused by two main strains of virus: equine-1 (H7N7) and equine-2 (H3N8). The OIE now cons ...
, equine infectious anemia
Equine infectious anemia or equine infectious anaemia (EIA), also known by horsemen as swamp fever, is a horse disease caused by a retrovirus (''Equine infectious anemia virus'') and transmitted by bloodsucking insects. The virus (''EIAV'') is en ...
and trypanosomiasis
Trypanosomiasis or trypanosomosis is the name of several diseases in vertebrates caused by parasitic protozoan trypanosomes of the genus ''Trypanosoma''. In humans this includes African trypanosomiasis and Chagas disease. A number of other diseas ...
. In addition, zebra hair is about as long as the mouthparts of these flies. Caro and colleagues (2019) reported this hypothesis as the "emerging consensus among biologists". It was found that flies preferred landing on solidly coloured surfaces over those with black-and-white striped patterns in 1930 by biologist R. Harris. A 2012 study concurred this and concluded that the stripes reflect contrasting light patterns rather than the more consistent patterns these insects use to locate food and habitat. A 2014 study found a correlation between striping and overlap with horse and tsetse fly
Tsetse ( , or ) (sometimes spelled tzetze; also known as tik-tik flies), are large, biting flies that inhabit much of tropical Africa. Tsetse flies include all the species in the genus ''Glossina'', which are placed in their own family, Glo ...
populations and activity. Other studies have found that zebras are rarely targeted by these insect species. Caro and colleagues studied captive zebras and horses and found that neither could deter flies from a distance, but zebra stripes made it difficult for flies to make a landing, both for zebras and horses dressed in zebra print coats. A 2020 study found that zebra stripes do not dazzle or work like a barber pole against flies since checkered patterns also repel them. White or light stripes painted on dark bodies have also been found to reduce fly irritations in both cattle and humans. In addition to the stripes, zebra can also defend against flies by swatting their tails.
Ecology and behaviour
Zebras may travel or migrate to wetter areas during the dry season. Plains zebras have been recorded travelling between Namibia and Botswana, the longest land migration of mammals in Africa. When migrating, they appear to rely on some memory of the locations where foraging conditions were best and may predict conditions months after their arrival. Plains zebras are more water-dependent and live in moister environments than other species. They usually can be found from a water source. Grévy's zebras can survive almost a week without water but will drink it every day when given the chance, and their bodies maintain water better than cattle. Mountain zebras can be found at elevations of up to . Zebras sleep for seven hours a day, standing up during the day and lying down during the night. They regularly use various objects as rubbing posts and will roll on the ground.
A zebra's diet is mostly grass
Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
es and sedge
The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus ''Carex'' wit ...
s, but they will opportunistically consume bark, leaves, buds, fruits, and roots. Compared to ruminants
Ruminants ( suborder Ruminantia) are hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microbial actions. The ...
, zebras have a simpler and less efficient digestive system. Nevertheless, they can subsist on lower-quality vegetation. Zebras may spend 60–80% of their time feeding, depending on the availability of vegetation. The plains zebra is a pioneer grazer, mowing down the upper, less nutritious grass canopy and preparing the way for more specialised grazers, which depend on shorter and more nutritious grasses below.
Zebras are preyed on mainly by lions. Leopards
The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, ...
, cheetahs
The cheetah (''Acinonyx jubatus'') is a large cat native to Africa and central Iran. It is the fastest land animal, estimated to be capable of running at with the fastest reliably recorded speeds being , and as such has evolved specialized ...
, spotted hyenas
The spotted hyena (''Crocuta crocuta''), also known as the laughing hyena, is a hyena species, currently classed as the sole extant member of the genus ''Crocuta'', native to sub-Saharan Africa. It is listed as being of least concern by the IUC ...
, brown hyena
The brown hyena (''Parahyaena brunnea''), also called strandwolf, is a species of hyena found in Namibia, Botswana, western and southern Zimbabwe, southern Mozambique and South Africa. It is the only extant species in the genus ''Parahyaena''. It ...
s and wild dogs
Wild Dogs is an American heavy metal band from Portland, Oregon, formed loosely in 1981 by original members Jeff Mark, Danny Kurth, Matt McCourt, and Pete Holmes. The band has since released six albums, three live albums and one DVD. Wild Dogs ...
pose less of a threat to adults. Nile crocodile
The Nile crocodile (''Crocodylus niloticus'') is a large crocodilian native to freshwater habitats in Africa, where it is present in 26 countries. It is widely distributed throughout sub-Saharan Africa, occurring mostly in the central, eastern ...
s also prey on zebras when they near water. Biting and kicking are a zebra's defense tactics. When threatened by lions, zebras flee, and when caught they are rarely effective in fighting off the big cats. In one study, the maximum speed of a zebra was found to be while a lion was measured at . Zebras do not escape lions by speed alone but by sideways turning, especially when the cat is close behind. With smaller predators like hyenas and dogs, zebras may act more aggressively, especially in defense of their young.
Social structure
Zebra species have two basic social structures. Plains and mountain zebras live in stable, closed family groups or harems
Harem (Persian: حرمسرا ''haramsarā'', ar, حَرِيمٌ ''ḥarīm'', "a sacred inviolable place; harem; female members of the family") refers to domestic spaces that are reserved for the women of the house in a Muslim family. A hare ...
consisting of one stallion
A stallion is a male horse that has not been gelded (castrated).
Stallions follow the conformation and phenotype of their breed, but within that standard, the presence of hormones such as testosterone may give stallions a thicker, "cresty" nec ...
, several mares
A mare is an adult female horse or other equine. In most cases, a mare is a female horse over the age of three, and a filly is a female horse three and younger. In Thoroughbred horse racing, a mare is defined as a female horse more than four y ...
, and their offspring. These groups have their own home ranges
A home range is the area in which an animal lives and moves on a periodic basis. It is related to the concept of an animal's territory which is the area that is actively defended. The concept of a home range was introduced by W. H. Burt in 1943. He ...
, which overlap, and they tend to be nomadic. Stallions form and expand their harems by recruiting young mares from their natal (birth) harems. The stability of the group remains even when the family stallion is displaced. Plains zebras groups gather into large herds and may create temporarily stable subgroups within a herd, allowing individuals to interact with those outside their group. Females in harems can spend more time feeding, and gain protection both for them and their young. The females have a linear dominance hierarchy
In biology, a dominance hierarchy (formerly and colloquially called a pecking order) is a type of social hierarchy that arises when members of animal social animal , social groups interact, creating a ranking system. A dominant higher-ranking i ...
with the high-ranking females being the ones that have lived in the group longest. While traveling, the most dominant females and their offspring lead the group, followed by the next most dominant. The family stallion trails behind. Young of both sexes leave their natal groups as they mature; females are usually herded by outside males to become part of their harems.
In the more arid-living Grévy's zebras, adults have more fluid associations and adult males establish large territories
A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal.
In international politics, a territory is usually either the total area from which a state may extract power resources or an ...
, marked by dung piles, and mate with the females that enter them. Grazing and drinking areas tend to be separated in these environments and the most dominant males establish territories near watering holes, which attract females with dependent foals and those who simply want a drink, while less dominant males control territories away from water with more vegetation, and only attract mares without foals. Mares may travel through several territories but remain in one when they have young. Staying in a territory offers a female protection from harassment by outside males, as well as access to resources.
In all species, excess males gather in bachelor group A bachelor herd is a herd of (usually) juvenile male animals who are still sexually immature or 'harem'-forming animals who have been thrown out of their parent groups but not yet formed a new family group. It may also refer to a group of males who ...
s. These are typically young males that are not yet ready to establish a harem or territory. With the plains zebra, the oldest males are the most dominant and group membership is stable. Bachelor groups tend to be at the boundaries of herds and during group movements, the bachelors trail behind or occupy the sides. Mountain zebra bachelor groups may also include young females that have left their natal group early, as well as old, former harem males. A territorial Grévy's zebra stallion may allow non-territorial bachelors in their territory, however when a mare in oestrous
The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous p ...
is present the territorial stallion keeps other stallions at bay. Bachelors prepare for their future harem roles with play fights and greeting/challenge rituals, which make up most of their activities.
Fights between males usually occur over mates and involve biting and kicking. In plains zebra, stallions fight each other over recently matured mares to bring into their group and her family stallion will fight off other males trying to abduct her. As long as a harem stallion is healthy, he is not usually challenged. Only unhealthy stallions have their harems taken over, and even then, the new stallion slowly takes over, peacefully displacing the old one. Agonistic behaviour
Agonistic behaviour is any social behaviour related to fighting. The term has broader meaning than aggressive behaviour because it includes threats, displays, retreats, placation, and conciliation. The term "agonistic behaviour" was first implemen ...
between male Grévy's zebras occurs at the border of their territories.
Communication
Zebras produce a number of vocalisations and noises. The plains zebra has a distinctive, barking contact call heard as "a-''ha'', a-''ha'', a-''ha''" or "kwa-ha, kaw-ha, ha, ha". The mountain zebra may produce a similar sound while the call of the Grévy's zebra has been described as "something like a hippo
The hippopotamus ( ; : hippopotamuses or hippopotami; ''Hippopotamus amphibius''), also called the hippo, common hippopotamus, or river hippopotamus, is a large semiaquatic mammal native to sub-Saharan Africa. It is one of only two extant ...
's grunt combined with a donkey's wheeze". Loud snorting and rough "gasping" in zebras signals alarm. Squealing is usually made when in pain, but can also be heard in friendly interactions. Zebras also communicate with visual displays, and the flexibility of their lips allows them to make complex facial expressions. Visual displays also consist of head, ear, and tail postures. A zebra may signal an intention to kick by dropping back its ears and whipping its tail. Flattened ears, bared teeth and a waving head may be used as threatening gestures by stallions.
Individuals may greet each other by rubbing and sniffing and then mutually rub their cheeks, and move along their bodies towards each other's genitals to sniff. They then may caress their shoulders against each other and lay their heads on one another. This greeting usually occurs between harem or territorial males or among bachelor males playing. Plains and mountain zebras strengthen their social bonds with grooming. Members of a harem nibble and rake along the neck, shoulder, and back with their teeth and lips. Grooming usually occurs between mothers and foals and between stallions and mares. Grooming establishes social rank and eases aggressive behaviour. Although Grévy's zebras generally do not perform social grooming.
Reproduction and parenting
Among plains and mountain zebras, the adult females mate only with their harem stallion, while in Grévy's zebras, mating is more promiscuous
Promiscuity is the practice of engaging in sexual activity frequently with different Sexual partner, partners or being indiscriminate in the choice of sexual partners. The term can carry a moral judgment. A common example of behavior viewed as pro ...
and the males have larger testes for sperm competition
Sperm competition is the competitive process between spermatozoa of two or more different males to fertilize the same egg during sexual reproduction. Competition can occur when females have multiple potential mating partners. Greater choice and ...
. Female zebras have five to ten day long oestrous cycles; physical signs include a swollen, everted (inside out) labia and copious flows of urine and mucus. Among reaching peak oestrous, mares spread-out their legs, lift their tails and open their mouths when in the presence of a male. Males assess the female's reproductive state with a curled lip and bared teeth (flehmen response
The flehmen response (; from German ''flehmen'', to bare the upper teeth, and Upper Saxon German ''flemmen'', to look spiteful), also called the flehmen position, flehmen reaction, flehmen grimace, flehming, or flehmening, is a behavior in which ...
) and the female will solicit mating by backing in. Gestation is typically around a year. A few days to a month later, mares can return to oestrus. In harem-holding species, oestrus in a female becomes less noticeable to outside males as she gets older, hence competition for older females is virtually nonexistent.
Usually, a single foal is born, which is capable of running within an hour of birth. A newborn zebra will follow anything that moves, so new mothers prevent other mares from approaching their foals as they become more familiar with the mother's striping pattern, smell and voice. At a few weeks old, foals begin to graze, but may continue to nurse for eight to thirteen months. Living in an arid environment, Grévy's zebras have longer nursing intervals and young only begin to drink water three months after birth.
In plains and mountain zebras, foals are cared for mostly by their mothers, but if threatened by pack-hunting hyenas and dogs, the entire group works together to protect all the young. The group forms a protective front with the foals in the centre, and the stallion will rush at predators that come too close. In Grévy's zebras, young stay in "kindergartens
Kindergarten is a preschool educational approach based on playing, singing, practical activities such as drawing, and social interaction as part of the transition from home to school. Such institutions were originally made in the late 18th cent ...
" when their mothers leave for water. These groups are tended to by the territorial male. A stallion may look after a foal in his territory to ensure that the mother stays, though it may not be his. By contrast, plains zebra stallions are generally intolerant of foals that are not theirs and may practice infanticide
Infanticide (or infant homicide) is the intentional killing of infants or offspring. Infanticide was a widespread practice throughout human history that was mainly used to dispose of unwanted children, its main purpose is the prevention of reso ...
and feticide
Foeticide (British English), or feticide (American and Canadian English), is the act of killing a fetus, or causing a miscarriage.
Etymology
Foeticide derives from two constituent Latin roots. ''Foetus'', meaning child, is an alternate form of ...
via violence to the pregnant mare.
Human relations
Cultural significance
With their distinctive black-and-white stripes, zebras are among the most recognisable mammals. They have been associated with beauty and grace, with naturalist Thomas Pennant
Thomas Pennant (14 June Old Style, OS 172616 December 1798) was a Welsh natural history, naturalist, traveller, writer and antiquarian. He was born and lived his whole life at his family estate, Downing Hall near Whitford, Flintshire, in Wales ...
describing them in 1781 as "the most elegant of quadrupeds". Zebras have been popular in photography, with some wildlife photographers describing them as the most photogenic animal. They have become staples in children's stories and wildlife-themed art, such as depictions of Noah's Ark
Noah's Ark ( he, תיבת נח; Biblical Hebrew: ''Tevat Noaḥ'')The word "ark" in modern English comes from Old English ''aerca'', meaning a chest or box. (See Cresswell 2010, p.22) The Hebrew word for the vessel, ''teva'', occurs twice in t ...
. In children's alphabet book
An alphabet book is a type of children's book giving basic instruction in an alphabet. Intended for young children, alphabet books commonly use pictures, simple language and alliteration to aid language learning. Alphabet books are published ...
s, the animals are often used to represent the letter 'Z'. Zebra stripes are also popularly used for body paintings, dress, furniture and architecture.
Zebras have been featured in African art
African art describes the modern and historical paintings, sculptures, installations, and other visual culture from native or indigenous Africans and the African continent. The definition may also include the art of the African diasporas, su ...
and culture
Culture () is an umbrella term which encompasses the social behavior, institutions, and norms found in human societies, as well as the knowledge, beliefs, arts, laws, customs, capabilities, and habits of the individuals in these groups.Tyl ...
for millennia. They are depicted in rock art
In archaeology, rock art is human-made markings placed on natural surfaces, typically vertical stone surfaces. A high proportion of surviving historic and prehistoric rock art is found in caves or partly enclosed rock shelters; this type also ...
in Southern Africa dating from 28,000 to 20,000 years ago, though less often than antelope species like eland. How the zebra got its stripes has been the subject of folk tales, some of which involve it being scorched by fire. The Maasai Maasai may refer to:
* Maasai people
*Maasai language
* Maasai mythology
* MAASAI (band)
See also
* Masai (disambiguation)
* Massai
Massai (also known as: Masai, Massey, Massi, Mah–sii, Massa, Wasse, Wassil or by the nickname "Big Foot" Mas ...
proverb "a man without culture is like a zebra without stripes" has become popular in Africa. The San people
The San peoples (also Saan), or Bushmen, are members of various Khoe, Tuu, or Kxʼa-speaking indigenous hunter-gatherer cultures that are the first cultures of Southern Africa, and whose territories span Botswana, Namibia, Angola, Zambia, ...
connected zebra stripes with water, rain and lighting, and water spirits were conceived of having zebra stripes.
For the Shona people
The Shona people () are part of the Bantu ethnic group native to Southern Africa, primarily living in Zimbabwe where they form the majority of the population, as well as Mozambique, South Africa, and a worldwide diaspora including global celebriti ...
, the zebra is a totem
A totem (from oj, ᑑᑌᒼ, italics=no or ''doodem'') is a spirit being, sacred object, or symbol that serves as an emblem of a group of people, such as a family, clan, lineage, or tribe, such as in the Anishinaabe clan system.
While ''the wo ...
animal and is glorified in a poem as an "iridescent and glittering creature". Its stripes have symbolised the union of male and female and at the ruined city of Great Zimbabwe
Great Zimbabwe is a medieval city in the south-eastern hills of Zimbabwe near Lake Mutirikwi and the town of Masvingo. It is thought to have been the capital of a great kingdom during the country's Late Iron Age about which little is known. Con ...
, zebra stripes decorate what is believed to be a ''domba'', a school meant to prepare girls for adulthood. In the Shona language
Shona (; sn, chiShona) is a Bantu language of the Shona people of Zimbabwe. It was codified by the colonial government in the 1950s. According to ''Ethnologue'', Shona, comprising the Zezuru, Korekore and Karanga dialects, is spoken by about 7 ...
, the name ''madhuve'' means "woman/women of the zebra totem" and is a name for girls in Zimbabwe
Zimbabwe (), officially the Republic of Zimbabwe, is a landlocked country located in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Botswana to the south-west, Zambia to the north, and Mozam ...
. The plains zebra is the national animal
This is a list of countries that have officially designated one or more animals as their national animals.
National animal
{, class="wikitable sortable"
! Country
! Name of animal
! Scientific name (Latin name)
! class="unsortable", Picture
...
of Botswana and zebras have been depicted on stamps during colonial
Colonial or The Colonial may refer to:
* Colonial, of, relating to, or characteristic of a colony or colony (biology)
Architecture
* American colonial architecture
* French Colonial
* Spanish Colonial architecture
Automobiles
* Colonial (1920 au ...
and post-colonial Africa. For people of the African diaspora
The African diaspora is the worldwide collection of communities descended from native Africans or people from Africa, predominantly in the Americas. The term most commonly refers to the descendants of the West and Central Africans who were e ...
, the zebra represented the politics of race and identity, being both black and white.
In cultures outside of its range, the zebra has been thought of as a more exotic alternative to the horse; the comic book character Sheena, Queen of the Jungle
Sheena, Queen of the Jungle, is a fictional American comic book jungle girl heroine, originally published primarily by Fiction House during the Golden Age of Comic Books. She was the first female comic book character with her own title, with her ...
, is depicted riding a zebra and explorer Osa Johnson
Martin Elmer Johnson (October 9, 1884 – January 13, 1937) and Osa Helen Johnson (née Leighty, March 14, 1894 – January 7, 1953) were married American adventurers and documentary filmmakers. In the first half of the 20th century the coupl ...
was photographed riding one. The film ''Racing Stripes
''Racing Stripes'' is a 2005 American sports comedy family film directed by Frederik Du Chau. The film was produced by Andrew A. Kosove, Broderick Johnson, Lloyd Phillips and Edward L. McDonnell, based on a script written by David Schmidt, Stev ...
'' features a captive zebra ostracised from the horses and ends up being ridden by a rebellious girl. Zebras have been featured as characters in animated films like ''Khumba
''Khumba'' is a 2013 South African computer-animated comedy film directed and co-produced by Anthony Silverston and written by Silverston and Raffaella Delle Donne. The film stars the voices of Jake T. Austin, Steve Buscemi, Loretta Devine, Lau ...
'', ''The Lion King
''The Lion King'' is a 1994 American animated musical drama film produced by Walt Disney Feature Animation and released by Walt Disney Pictures. The 32nd Disney animated feature film and the fifth produced during the Disney Renaissance, it ...
'' and the ''Madagascar'' films and television series such as '' Zou''.
Zebras have been popular subjects for abstract, modernist
Modernism is both a philosophical and arts movement that arose from broad transformations in Western society during the late 19th and early 20th centuries. The movement reflected a desire for the creation of new forms of art, philosophy, an ...
and surrealist
Surrealism is a cultural movement that developed in Europe in the aftermath of World War I in which artists depicted unnerving, illogical scenes and developed techniques to allow the unconscious mind to express itself. Its aim was, according to l ...
artists. Such art includes Christopher Wood's ''Zebra and Parachute'', Lucian Freud
Lucian Michael Freud (; 8 December 1922 – 20 July 2011) was a British painter and draughtsman, specialising in figurative art, and is known as one of the foremost 20th-century English portraitists. He was born in Berlin, the son of Jewis ...
's ''The Painter's Room'' and ''Quince on a Blue Table'' and the various paintings of Mary Fedden
Mary Fedden, OBE RA RWA (14 August 1915 – 22 June 2012) was a British artist.
Early years
Sometimes mistakenly described as the daughter of Roy Fedden (who was in fact her uncle, as was Romilly Fedden), Mary Fedden was born in Bristol ...
and Sidney Nolan
Sir Sidney Robert Nolan (22 April 191728 November 1992) was one of Australia's leading artists of the 20th century. Working in a wide variety of mediums, his oeuvre is among the most diverse and prolific in all of modern art. He is best known ...
. Victor Vasarely
Victor Vasarely (; born Győző Vásárhelyi, ; 9 April 1906 – 15 March 1997) was a Hungarian-French artist, who is widely accepted as a "grandfather" and leader of the Op art movement.
His work entitled ''Zebra'', created in 1937, is consi ...
depicted zebras as black and white lines and connected in a jigsaw puzzle
A jigsaw puzzle is a tiling puzzle that requires the assembly of often irregularly shaped interlocking and mosaiced pieces, each of which typically has a portion of a picture. When assembled, the puzzle pieces produce a complete picture.
In th ...
fashion. Carel Weight
Carel Victor Morlais Weight, (10 September 1908 – 13 August 1997) was an English painter.
Biography
Weight was born in Paddington in 1908. His father was a bank cashier and his mother, who was of Swedish and German descent, was a chirop ...
's ''Escape of the Zebra from the Zoo during an Air Raid'' was based on a real life incident of a zebra escaping during the bombing of London Zoo and consists of four comic book-like panels. Zebras have lent themselves to products and advertisements, notably for 'Zebra Grate Polish' cleaning supplies by British manufacturer Reckitt and Sons
Reckitt and Sons was a leading British manufacturer of household products, notably starch, black lead, laundry blue, and household polish, and based in Kingston upon Hull.
Isaac Reckitt began business in Hull in 1840, and his business becam ...
and Japanese pen manufacturer Zebra Co., Ltd.
Captivity
Zebras have been kept in captivity since at least the Roman Empire
The Roman Empire ( la, Imperium Romanum ; grc-gre, Βασιλεία τῶν Ῥωμαίων, Basileía tôn Rhōmaíōn) was the post-Republican period of ancient Rome. As a polity, it included large territorial holdings around the Mediterr ...
. In later times, captive zebras have been shipped around the world, often for diplomatic reasons. In 1261, Sultan Baibars
Al-Malik al-Zahir Rukn al-Din Baybars al-Bunduqdari ( ar, الملك الظاهر ركن الدين بيبرس البندقداري, ''al-Malik al-Ẓāhir Rukn al-Dīn Baybars al-Bunduqdārī'') (1223/1228 – 1 July 1277), of Turkic Kipchak ...
of Egypt
Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Mediter ...
established an embassy with Alfonso X of Castile
Alfonso X (also known as the Wise, es, el Sabio; 23 November 1221 – 4 April 1284) was King of Castile, León and Galicia from 30 May 1252 until his death in 1284. During the election of 1257, a dissident faction chose him to be king of Germ ...
and sent a zebra and other exotic animals as gifts. In 1417, a zebra was gifted to the Chinese people by Somalia and displayed before the Yongle Emperor
The Yongle Emperor (; pronounced ; 2 May 1360 – 12 August 1424), personal name Zhu Di (), was the third Emperor of the Ming dynasty, reigning from 1402 to 1424.
Zhu Di was the fourth son of the Hongwu Emperor, the founder of the Ming dyn ...
. The fourth Mughal emperor Jahangir
Nur-ud-Din Muhammad Salim (30 August 1569 – 28 October 1627), known by his imperial name Jahangir (; ), was the fourth Mughal Emperor, who ruled from 1605 until he died in 1627. He was named after the Indian Sufi saint, Salim Chishti.
Ear ...
received a zebra from Ethiopia in 1620 and Ustad Mansur
Ustad Mansur (flourished 1590-1624) was a seventeenth-century Mughal painter and court artist. He grew up during the reign of Jahangir (r. 1605 - 1627) during which period he excelled at depicting plants and animals. He was the earliest artist t ...
made a painting of it. In the 1670s, Ethiopian Emperor
The emperor of Ethiopia ( gez, ንጉሠ ነገሥት, nəgusä nägäst, " King of Kings"), also known as the Atse ( am, ዐፄ, "emperor"), was the hereditary ruler of the Ethiopian Empire, from at least the 13th century until the abolition ...
Yohannes I
Yohannes I ( gez, ቀዳማዊ ዮሐንስ), also known as Yohannes the Righteous (Ge'ez: ጻድቁ ዮሐንስ), throne name A'ilaf Sagad (Ge'ez: አእላፍ ሰገድ; 1640 – 19 July 1682) was Emperor of Ethiopia from 1667 to 1682, and ...
exported two zebras to the Dutch governor of Jakarta
Jakarta (; , bew, Jakarte), officially the Special Capital Region of Jakarta ( id, Daerah Khusus Ibukota Jakarta) is the capital and largest city of Indonesia. Lying on the northwest coast of Java, the world's most populous island, Jakarta ...
. These animals would eventually be given by the Dutch to the Tokugawa Shogunate
The Tokugawa shogunate (, Japanese 徳川幕府 ''Tokugawa bakufu''), also known as the , was the military government of Japan during the Edo period from 1603 to 1868. Nussbaum, Louis-Frédéric. (2005)"''Tokugawa-jidai''"in ''Japan Encyclopedia ...
of Japan.
When Queen Charlotte
Charlotte of Mecklenburg-Strelitz (Sophia Charlotte; 19 May 1744 – 17 November 1818) was Queen of Great Britain and of Ireland as the wife of King George III from their marriage on 8 September 1761 until the union of the two kingdoms ...
received a zebra as a wedding gift in 1762, the animal became a source of fascination for the people of Britain. Many flocked to see it at its paddock
A paddock is a small enclosure for horses. In the United Kingdom, this term also applies to a field for a general automobile racing competition, particularly Formula 1.
Description
In Canada and the United States of America, a paddock is a small ...
at Buckingham Palace
Buckingham Palace () is a London royal residence and the administrative headquarters of the monarch of the United Kingdom. Located in the City of Westminster, the palace is often at the centre of state occasions and royal hospitality. It ...
. It soon became the subject of humour and satire, being referred to as "The Queen's Ass", and was the subject of an oil painting by George Stubbs
George Stubbs (25 August 1724 – 10 July 1806) was an English painter, best known for his paintings of horses. Self-trained, Stubbs learnt his skills independently from other great artists of the 18th century such as Reynolds or Gainsborough ...
in 1763. The zebra also gained a reputation for being ill-tempered and kicked at visitors. In 1882, Ethiopia sent a zebra to French president Jules Grévy
François Judith Paul Grévy (15 August 1807 – 9 September 1891), known as Jules Grévy (), was a French lawyer and politician who served as President of France from 1879 to 1887. He was a leader of the Moderate Republicans, and given that hi ...
, and the species it belonged to was named in his honour.
Attempts to domesticate
Domestication is a sustained multi-generational relationship in which humans assume a significant degree of control over the reproduction and care of another group of organisms to secure a more predictable supply of resources from that group. A ...
zebras were largely unsuccessful. It is possible that having evolved under pressure from the many large predators of Africa, including early humans, they became more aggressive, thus making domestication more difficult. However, zebras have been trained
Training is teaching, or developing in oneself or others, any skills and knowledge or fitness that relate to specific useful competencies. Training has specific goals of improving one's capability, capacity, productivity and performance. I ...
throughout history. In Rome, zebras are recorded to have pulled chariots during amphitheatre
An amphitheatre (British English) or amphitheater (American English; both ) is an open-air venue used for entertainment, performances, and sports. The term derives from the ancient Greek ('), from ('), meaning "on both sides" or "around" and ...
games starting in the reign of Caracalla
Marcus Aurelius Antoninus (born Lucius Septimius Bassianus, 4 April 188 – 8 April 217), better known by his nickname "Caracalla" () was Roman emperor from 198 to 217. He was a member of the Severan dynasty, the elder son of Emperor S ...
(198 to 217 AD). In the late 19th century, the zoologist Walter Rothschild
Lionel Walter Rothschild, 2nd Baron Rothschild, Baron de Rothschild, (8 February 1868 – 27 August 1937) was a British banker, politician, zoologist and soldier, who was a member of the Rothschild family. As a Zionist leader, he was present ...
trained some zebras to draw a carriage in England, which he drove to Buckingham Palace to demonstrate that it can be done. However, he did not ride on them knowing that they were too small and aggressive. In the early 20th century, German colonial officers in East Africa tried to use zebras for both driving and riding, with limited success.
Conservation
As of 2016–2019, the IUCN Red List
The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
of mammals lists the Grévy's zebra as endangered
An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inva ...
, the mountain zebra as vulnerable and the plains zebra as near-threatened
A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify fo ...
. Grévy's zebra populations are estimated at less than 2,000 mature individuals, but they are stable. Mountain zebras number near 35,000 individuals and their population appears to be increasing. Plains zebra are estimated to number 150,000–250,000 with a decreasing population trend. Human intervention has fragmented zebra ranges and populations. Zebras are threatened by hunting for their hide and meat, and habitat destruction
Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
. They also compete with livestock and have their travelling routes obstruct by fences. Civil wars in some countries have also caused declines in zebra populations. By the early 20th century, zebra skins were being used to make rugs and chairs. In the 21st century, zebras may be taken by trophy hunters as zebra skin rugs sell for $1,000 to $2,000. Trophy hunting was rare among African peoples though the San were known to hunt zebra for meat.
The quagga population was hunted by early Dutch settlers and later by Afrikaner
Afrikaners () are a South African ethnic group descended from Free Burghers, predominantly Dutch settlers first arriving at the Cape of Good Hope in the 17th and 18th centuries.Entry: Cape Colony. ''Encyclopædia Britannica Volume 4 Part 2: ...
s to provide meat or for their skins. The skins were traded or used locally. The quagga was probably vulnerable to extinction due to its restricted range, and because they were easy to find in large groups. The last known wild quagga died in 1878. The last captive quagga, a female in Amsterdam's Natura Artis Magistra
Natura Artis Magistra (Latin for "Nature is the teacher of the arts"), commonly known just as Artis (), is a zoo and botanical garden in the centre of Amsterdam. It is the oldest zoo in the Netherlands and fifth oldest zoo in the world.
In addit ...
zoo, lived there from 9 May 1867 until it died on 12 August 1883. The Cape mountain zebra
The Cape mountain zebra (''Equus zebra zebra'') is a subspecies of mountain zebra that occurs in certain mountainous regions of the Western and Eastern Cape provinces of South Africa.
It is the smallest of all existing zebra species and also the ...
, a subspecies of mountain zebra, nearly went extinct due to hunting and habitat destruction, with less than 50 individuals left by the 1950s. Protections from South African National Parks
South African National Parks (SANParks) is the body responsible for managing South Africa's national parks. SANParks was formed in 1926, and currently manages 19 parks consisting of , over 3% of the total area of South Africa.
Many parks offer ...
allowed the population to rise to 2,600 by the 2010s.
Zebras can be found in numerous protected areas. Important areas for the Grévy's zebra include Yabelo Wildlife Sanctuary
Yabelo Wildlife Sanctuary is a protected area and wildlife sanctuary in southern Ethiopia. It is located in the Borena Zone of the Oromia Region west of the town of Yabelo, having an area of 2,500 square kilometers and elevations ranging from 143 ...
and Chelbi Sanctuary in Ethiopia and Buffalo Springs, Samburu and Shaba National Reserves in Kenya
)
, national_anthem = "Ee Mungu Nguvu Yetu"()
, image_map =
, map_caption =
, image_map2 =
, capital = Nairobi
, coordinates =
, largest_city = Nairobi
, ...
. The plains zebra inhabitats the Serengeti National Park
The Serengeti National Park is a large national park in northern Tanzania that stretches over . It is located entirely in eastern Mara Region and north east portion of Simiyu Region and contains over of virgin savanna. The park was established in ...
in Tanzania, Tsavo
Tsavo is a region of Kenya located at the crossing of the Uganda Railway over the Tsavo River, close to where it meets the Athi-Galana-Sabaki River. Two national parks, Tsavo East and Tsavo West are located in the area.
The meaning of the wor ...
and Masai Mara
Maasai Mara, also sometimes spelled Masai Mara and locally known simply as The Mara, is a large national game reserve in Narok, Kenya, contiguous with the Serengeti National Park in Tanzania. It is named in honor of the Maasai people, the ancest ...
in Kenya, Hwange National Park
Hwange National Park (formerly Wankie Game Reserve) is the largest natural reserve in Zimbabwe. It is around 14,600 sq km in area. It lies in the northwest of the country, just off the main road between Bulawayo and Victoria Falls. The neares ...
in Zimbabwe, Etosha National Park
Etosha National Park is a national park in northwestern Namibia and one of the largest national parks in Africa. It was proclaimed a game reserve in March 1907 in Ordinance 88 by the Governor of German South West Africa, Friedrich von Lindequist. ...
in Namibia
Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
, and Kruger National Park
Kruger National Park is a South African National Park and one of the largest game reserves in Africa. It covers an area of in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends from north to south and from ea ...
in South Africa
South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
. Mountain zebras are protected in Mountain Zebra National Park
Mountain Zebra National Park is a national park in the Eastern Cape province of South Africa proclaimed in July 1937 for the purpose of providing a nature reserve for the endangered Cape mountain zebra.
History
In the early 1930s, the Cape mou ...
, Karoo National Park
The Karoo ( ; from the Afrikaans borrowing of the South Khoekhoe Khoe languages, !Orakobab or Khoemana word ''ǃ’Aukarob'' "Hardveld") is a semi-desert natural region of South Africa. No exact definition of what constitutes the Karoo is availa ...
and Goegap Nature Reserve
The Goegap Nature Reserve is a semi-desert nature reserve in Northern Cape, South Africa and lies around east of Springbok. The area was once used to graze livestock and was proclaimed in 1966 as a reserve then fenced in 1969 and wildlife introd ...
in South Africa as well as Etosha and Namib-Naukluft Park
The Namib-Naukluft Park is a national park in western Namibia, situated between the coast of the Atlantic Ocean and the edge of the Great Escarpment. It encompasses part of the Namib Desert (considered the world's oldest desert), the Naukluft mo ...
in Namibia.
See also
* Fauna of Africa
The fauna of Africa, in its broader sense, is all the animals living in Africa and its surrounding seas and islands. The more characteristic African fauna is found in the Afrotropical realm. Lying almost entirely within the tropics, and equally to ...
* Lord Morton's mare
Lord Morton’s mare was an equid hybrid and once an often-noticed example in the history of evolutionary theory.
In 1820, George Douglas, 16th Earl of Morton, F.R.S., reported to the President of the Royal Society that, being desirous of domes ...
* Primitive markings
Primitive markings are a group of hair coat markings and qualities seen in several equine species, including horses, donkeys, and asses. In horses, they are associated with primitive breeds, though not limited to such breeds. The markings are ...
– markings found on other equines
* Zonkey (Tijuana)
Zonkeys in Tijuana, Mexico are donkeys (also known as burros in Mexico and the Southwestern United States; ''burro'' is the Spanish word for a small donkey) painted with fake zebra stripes, so that tourists will pay the owner to appear in souveni ...
– a donkey painted with zebra stripes
Citations
General bibliography
*
*
External links
The Quagga Project
An organisation that selectively breeds zebras to recreate the hair coat pattern of the quagga
{{Featured article
*
Equus (genus)
Fauna of Sub-Saharan Africa
Herbivorous mammals
Mammal common names
Mammals of Africa