Zanthoxylum Rhetsa
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''Zanthoxylum rhetsa'', commonly known as Indian prickly ash, is a species of flowering plant in the family
Rutaceae The Rutaceae is a family, commonly known as the rueRUTACEAE
in BoDD – Botanical Derm ...
and occurs from India east to the Philippines and south to northern Australia. It is a deciduous shrub or tree with cone-shaped spines on the stems,
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
leaves with between nine and twenty-three leaflets,
panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...
s of white or yellowish, male and female flowers, followed by spherical red, brown or black follicles.


Description

''Zanthoxylum rhetsa'' is a shrub or tree that sometimes grows to a height of . The plant is sometimes
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
and has stems with thick, cone-shaped spines on the older stems. The leaves are long and pinnate, with nine to twenty three egg-shaped to elliptical leaflets. The leaflets are long and wide, the side leaflets on petiolules long and the end leaflet on a petiolule long. The flowers are arranged on the ends of branchlets, sometimes also in leaf axils, in panicles up to long. Each flower is on a
pedicel Pedicle or pedicel may refer to: Human anatomy *Pedicle of vertebral arch, the segment between the transverse process and the vertebral body, and is often used as a radiographic marker and entry point in vertebroplasty and kyphoplasty procedures ...
long, the four
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
s joined at the base and long and the four petals white or yellowish white and long. Male flowers have
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s about long with a sterile
carpel Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
about long. Female flowers lack stamens and usually have a single carpel about long. Flowering occurs in summer and the fruit is a spherical red or brown to black follicle wide.


Taxonomy

Indian prickly ash was first formally described in 1820 by
William Roxburgh William Roxburgh FRSE FRCPE Linnean Society of London, FLS (3/29 June 1751 – 18 February 1815) was a Scottish people, Scottish surgeon and botanist who worked extensively in India, describing species and working on economic botany. He is known ...
who gave it the name ''Fagara rhetsa'' in his book, ''Flora Indica''. In 1824,
de Candolle Augustin Pyramus (or Pyrame) de Candolle (, , ; 4 February 17789 September 1841) was a Swiss botanist. René Louiche Desfontaines launched de Candolle's botanical career by recommending him at a herbarium. Within a couple of years de Candolle ...
changed the name to ''Zanthoxylum rhetsa'' in his book ''
Prodromus Systematis Naturalis Regni Vegetabilis ''Prodromus Systematis Naturalis Regni Vegetabilis'' (1824–1873), also known by its standard botanical abbreviation ''Prodr. (DC.)'', is a 17-volume treatise on botany initiated by Augustin Pyramus de Candolle. De Candolle intended it as a summa ...
''.


Distribution and habitat

''Zanthoxylum rhetsa'' grows in rainforest and coastal thickets from sea level to an altitude of , and is found in India, east to the Philippines and south to northern Australia. It occurs in the northern Kimberley in Western Australia, the northern coastal areas of the Northern Territory,
Cape York Peninsula Cape York Peninsula is a large peninsula located in Far North Queensland, Australia. It is the largest unspoiled wilderness in northern Australia.Mittermeier, R.E. et al. (2002). Wilderness: Earth’s last wild places. Mexico City: Agrupación ...
in Queensland and on Gabba and Moa Islands in the
Torres Strait The Torres Strait (), also known as Zenadh Kes, is a strait between Australia and the Melanesian island of New Guinea. It is wide at its narrowest extent. To the south is Cape York Peninsula, the northernmost extremity of the Australian mai ...
.


Uses

The people of
Goa Goa () is a state on the southwestern coast of India within the Konkan region, geographically separated from the Deccan highlands by the Western Ghats. It is located between the Indian states of Maharashtra to the north and Karnataka to the ...
, the
Konkan The Konkan ( kok, कोंकण) or Kokan () is a stretch of land by the western coast of India, running from Damaon in the north to Karwar in the south; with the Arabian Sea to the west and the Deccan plateau in the east. The hinterland ...
and
Kanara Kanara, also known as Karavali is the historically significant stretch of land situated by the southwestern coast of India, alongside the Arabian Sea in the present-day Indian state of Karnataka. The region comprises three civil districts, ...
coasts, and
Coorg Kodagu (also known by its former name Coorg) is an administrative district in the Karnataka state of India. Before 1956, it was an administratively separate Coorg State, at which point it was merged into an enlarged Mysore State. It occupies ...
use the woody pericarp of the tiny fruits as a spice, particularly with seafood dishes. The spice is known as "triphal" in Marathi and "teppal" in Konkani - both words referring to the three lobes of the pericarp. The spice contains a chemical ingredient,
sanshool Hydroxy-''alpha''-sanshool is a molecule found in plants from the genus ''Zanthoxylum''. It is believed to be responsible for the numbing and tingling sensation caused by eating food cooked with Sichuan peppercorns and Uzazi. The term ''sansho ...
, a local anesthetic that causes a tingling sensation on the tongue. Sanshool is also the main principle of Sichuan Pepper, which comes from the related species
Zanthoxylum bungeanum ''Zanthoxylum bungeanum '' is a species of plant in the family Rutaceae. It is one of the sources of the spice Sichuan pepper. The plant is native to North-Central China, South-Central China, Southeast China, East Himalayas The Himalayas ...
and the Japanese/Korean pepper
Zanthoxylum piperitum ''Zanthoxylum piperitum'', also known as Japanese pepper or Japanese prickly-ash is a deciduous aromatic spiny shrub or small tree of the citrus and rue family Rutaceae, native to Japan and Korea. It is called sanshō () in Japan and chopi () ...
. Many butterflies, including ''
Papilio buddha ''Papilio buddha'', the Malabar banded peacock, is a species of swallowtail butterfly found in the Western Ghats of India. The Government of Kerala declared it as the official Kerala state butterfly Description ''P. buddha'' resembles '' P ...
'' and ''
Papilio helenus ''Papilio helenus'', the red Helen, is a large swallowtail butterfly found in forests of southern India and parts of southeast Asia. Range ''Papilio helenus'' is rarely found in Sri Lanka, southern and north-east India, Nepal, Bhutan, Banglades ...
'', use this as a
host plant In biology and medicine, a host is a larger organism that harbours a smaller organism; whether a parasitic, a mutualistic, or a commensalist ''guest'' (symbiont). The guest is typically provided with nourishment and shelter. Examples include a ...
.


See also

* Sichuan pepper


References

{{Taxonbar, from=Q13114182 rhetsa Medicinal plants Plants described in 1814 Taxa named by William Roxburgh Flora of Queensland Rosids of Western Australia Flora of India (region) Flora of Southeast Asia