Yunnanocephalidae
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''Yunnanocephalus'' is a genus of ptychopariid trilobite. It lived during the late Atdabanian and Botomian stages, in what are currently Antarctica, Australia and China. It was a "moderately common" member of the Chengjiang Fauna. ''Yunnanocephalus'' is the only genus currently assigned to the Yunnanocephalidae family.


Description

''Yunnanocephalus'' is a small (about ) trilobite with an inverted elongated egg-shaped outline. Its headshield (or
cephalon Cephalon, Inc. was an American biopharmaceutical company co-founded in 1987 by pharmacologist Frank Baldino, Jr., neuroscientist Michael Lewis, and organic chemist James C. Kauer—all three former scientists with the DuPont Company. Baldino s ...
) is ovate and twice as wide as long, slightly wider than and not confluent with the articulate middle part of the exoskeleton (or thorax). The raised central section (or glabella) is weakly defined, without clear furrows except for one defining the occipital ring, tapering forward, with the front straight and confluent with the eye ridges. The occipital ring is about as wide as the border and the axial rings of the thorax. The distance between the glabella and the frontal border is slightly wider that the border. The visual surface of the eye is rather small (about 20% of the length of the cephalon), with the front about halflength of the cephalon. The natural rupture line (or suture) anterior of the visual surface is parallel to the midline or converges slightly forward and from the back of the eye diverges to the middle of the widely rounded back edges of the cephalon. The thorax has 14 segments, with short triangular spines, the 3rd and 4th slightly wider than the frontal segments, and a small tailshield (or pygidium) consisting almost fully of the axis and post axial boss, with two vaguely defined axial rings.


Ontogeny

During the ontogenetic development of ''Y. yunnanensis'' between a late larval phase ( meraspis degree 9, 2½mm) and the adult (or holaspis) the
facial suture Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...
changes from proparian to gonatoparian. Although this development has also been observed in '' Duyunaspis duyunensis'', it is really uncommon. Usually, proparian holaspid trilobites developed from
opisthoparian Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...
meraspids due to a mechanism called heterochrony.


Distribution

* ''Y. longioccipitalis'' is known from the Lower Cambrian of Antarctica (Atdabanian, Shackleton Limestone, 81.9° S, 158.7° E). * ''Y. macromelos'' has been found in the Lower Cambrian of Australia (Botomian, Mernmerna Formation, Angorichina station, east side of The Bunkers, 1 km S Balcoracana Creek, South-Australia, 31.2° S, 138.9° E). * ''Y. yunnanensis'' occurs in the Lower Cambrian of China (Atdabanian, Yuanshan or Qiongzhusi Formation,
Chengjiang biota The Maotianshan Shales are a series of Early Cambrian deposits in the Chiungchussu Formation, famous for their '' Konservat Lagerstätten'', deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales ...
, near Haikou, Anning, Jinning, Jingmacun, Kunming, Yunnan, 24.8° N, 102.6° E; and Maotianshan Bed 4, 24.0° N, 102.0° E). ''cited on''


References


External links


Order Pitychopariida at trilobite.info
{{Taxonbar, from=Q5974962 Ellipsocephaloidea Ptychopariida genera Cambrian trilobites Trilobites of Antarctica Cambrian trilobites of Australia Cambrian trilobites of Asia Fossils of China Fossil taxa described in 1936 Cambrian genus extinctions