Yunia Nakibande
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''Yunia'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of extinct
vascular plant Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They al ...
s from the Early Devonian (Pragian or Siegenian stage, around ). It was first described from the Posongchong Formation of
Yunnan Yunnan , () is a landlocked Provinces of China, province in Southwest China, the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is ...
, China. The leafless plant consisted of spiny stems, some 2 to 5 cm wide, which branched dichotomously at wide angles in a cruciate arrangement. Each stem contained vascular tissue with one or two strands of protoxylem. The spore-forming organs (
sporangia A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cy ...
) were elongated and borne on short stalks. The spores had a relatively smooth sculptural pattern and were trilete (i.e. each spore has three lines on it resulting from its formation in a tetrahedral set of four spores). In 2004, Crane et al. published a simplified cladogram for the polysporangiophytes in which ''Yunia'' is basal to the
lycophytes The lycophytes, when broadly circumscribed, are a vascular plant (tracheophyte) subgroup of the kingdom Plantae. They are sometimes placed in a division Lycopodiophyta or Lycophyta or in a subdivision Lycopodiophytina. They are one of the oldest ...
(clubmosses and relatives). It had previously been placed in the "
trimerophyte Trimerophytopsida (or Trimeropsida) is a class of early vascular plants from the Devonian, informally called trimerophytes. It contains genera such as ''Psilophyton''. This group is probably paraphyletic, and is believed to be the ancestral group ...
s" (a group now thought to be paraphyletic), which were considered to have given rise to all the other vascular plants ''except'' the lycophytes. Hao and Xue in 2013 considered the genus as a questionable zosterophyll.


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Cladogram
from {{Taxonbar, from=Q8061285 Early Devonian plants Zosterophylls Prehistoric lycophyte genera