Ypsolopha Ustella Two Leaves Of Hornbeam Fastened Together By Larva
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''Ypsolopha'' is a genus of
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s of the family
Ypsolophidae Ypsolophidae is a family of moths with some 160 species. They are included in the Plutellidae by many authors. Systematics This is a list of genera and selected species. Ypsolophinae * '' Phrealcia'' Chrétien, 1900 * '' Ypsolopha'' Latreille, ...
. It is the type genus of the family and comprises over 120 described species (about 95% of the family's known world diversity).


Distribution

Most ''Ypsolopha'' species have been recorded from the
Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical region ...
temperate region.


Description

''Ypsolopha'' species are variable in shape and color and no exclusive superficial features have been established for the group. In contrast, the genitalia of both sexes are remarkably homogeneous.


Biology

Adults are nocturnal or rarely diurnal. Their resting postures are various, but they often have the head down and the lower body up. '' Ypsolopha acuminata'' mimics a small broken branch at rest. The
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e usually live in open webs on the leaves of various, primarily woody, plants and mostly feed on a limited range of host plants. They are active primarily at night and have two defensive behaviors that involve wiggling and jumping.


Selected species

*'' Ypsolopha acerella'' Ponomarenko, Sohn & Zinchenko, 2011 *'' Ypsolopha acuminata'' (Butler, 1878) *''
Ypsolopha albiramella ''Ypsolopha albiramella'' is a moth of the family Ypsolophidae. It is known from Italy, Croatia, Albania, North Macedonia, Greece, Crete and Turkmenistan. The larvae have been recorded feeding on ''Ephedra distachya ''Ephedra distachya'' is a ...
'' (Mann, 1861) *'' Ypsolopha albistriatus'' (Issiki, 1930) *''
Ypsolopha aleutianella ''Ypsolopha aleutianella'' is a moth of the family Ypsolophidae. It is known from the Aleutian Islands The Aleutian Islands (; ; ale, Unangam Tanangin,”Land of the Aleuts", possibly from Chukchi language, Chukchi ''aliat'', "island"), also ...
'' (Beutenmüller, 1889) *''
Ypsolopha alpella ''Ypsolopha alpella'' is a moth of the family Ypsolophidae. It is found in southern and central Europe and Siberia. The wingspan is approximately 16 mm. The moth flies from June to October depending on the location. The larvae feed on oak ...
'' (Denis & Schiffermüller, 1775) *'' Ypsolopha amoenella'' (Christoph, 1882) *'' Ypsolopha angelicella'' (Busck, 1903) *''
Ypsolopha arizonella ''Ypsolopha arizonella'' is a moth of the family Ypsolophidae. It is known from the United States, including Arizona. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. F ...
'' (Busck, 1903) *''
Ypsolopha asperella ''Ypsolopha asperella'' is a moth of the family Ypsolophidae. It is found in Northern and Central Europe, Siberia, Korea, China, Asia Minor and Mideast Asia. The wingspan is 20–21 mm. The larvae feed on apple, ''Crataegus'', '' Cerasus'' ...
'' (Linnaeus, 1761) *'' Ypsolopha atrobrunnella'' Ponomarenko & Sohn, 2011Ponomarenko, M.G., Sohn, J.-C., Zinchenko, Y.N., & Wu, C.-S. "Five new East-Asian species of the genus ''Ypsolopha'' Latreille (Lepidoptera: Ypsolophidae)." ''Zootaxa'' 2760 (2011): 18-28. *'' Ypsolopha aurata'' Moriuti, 1977 *'' Ypsolopha barberella'' (Busck, 1903) *'' Ypsolopha blandella'' (Christoph, 1882) *''
Ypsolopha buscki ''Ypsolopha buscki'' is a moth of the family Ypsolophidae. It is known from the United States, including Arizona. The wingspan is about 22 mm. The antennae are light fuscous. The labial palpi are blackish fuscous on the outside and light o ...
'' Heppner, 1982 *'' Ypsolopha cajaliella'' Vives, 2003 *'' Ypsolopha canariella'' (Walsingham, 1881) *''
Ypsolopha cervella ''Ypsolopha cervella'' is a moth of the family Ypsolophidae. It is found from Vancouver Island southward along the Pacific coast to San Diego County in California and inland to Arizona. The wingspan is about 19 mm. There is one generation p ...
'' (Walsingham, 1881) *''
Ypsolopha chazariella ''Ypsolopha chazariella'' is a moth of the family Ypsolophidae. It is known from Latvia, Lithuania, the Czech Republic, Slovakia, Albania, Bosnia and Herzegovina, Hungary, Bulgaria, Romania and France. It has also been recorded from Finland ...
'' (Mann, 1866) *''
Ypsolopha cockerella ''Ypsolopha cockerella'' is a moth of the family Ypsolophidae. It is known from the United States, including New Mexico. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip ...
'' (Busck, 1903) *''
Ypsolopha colleaguella ''Ypsolopha colleaguella'' is a moth of the family Ypsolophidae. It is known from the Volga River valley in southern Russia and Kazakhstan (the area of the lower Irtysz River near the Chinese border). The wingspan is 17.5–19.5 mm. The fo ...
'' Baraniak, 2007 *''
Ypsolopha contractella ''Ypsolopha contractella'' is a moth of the family Ypsolophidae. It is known from Japan (the islands of Hokkaido and Honshu), Korea, north-eastern China and the Russian Far East The Russian Far East (russian: Дальний Восток Р ...
'' (Caradja, 1920) *''
Ypsolopha coriacella ''Ypsolopha coriacella'' is a moth of the family Ypsolophidae. It is known from Germany, Poland, Austria, Switzerland, Slovakia, the Czech Republic and Russia. The larvae feed on ''Abies'' species. However, the species occurs in the regions with ...
'' (Herrich-Schäffer, 1855) *'' Ypsolopha costibasella'' (Caradja, 1939) *'' Ypsolopha cristata'' Moriuti, 1977 *''
Ypsolopha delicatella ''Ypsolopha delicatella'' is a moth of the family Ypsolophidae. It is known from the United States, including Arizona and California. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the ...
'' (Busck, 1903) *''
Ypsolopha dentella ''Ypsolopha dentella'', the honeysuckle moth, is a moth of the family Ypsolophidae. It is found in Europe, Anatolia, north-eastern China, Russia and mideast Asia. It is also present in North America, where it is known from the eastern United Stat ...
'' (Fabricius, 1775) *'' Ypsolopha dentiferella'' (Walsingham, 1881) *''
Ypsolopha diana ''Ypsolopha diana'' is a moth of the family Ypsolophidae. It is known from China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 ...
'' (Caradja, 1939) *''
Ypsolopha distinctatus ''Ypsolopha distinctata'' is a moth of the family Ypsolophidae. It is known from Japan. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777– ...
'' Moriuti, 1977 *''
Ypsolopha divisella ''Ypsolopha divisella'' is a moth of the family Ypsolophidae. It is from France. The wingspan is 16–17 mm. The larvae feed ''Ephedra altissima ''Ephedra altissima'' is a species of ''Ephedra'' that is native to the western Sahara (Mo ...
'' (Chrétien, 1915) *''
Ypsolopha dorsimaculella ''Ypsolopha dorsimaculella'' is a moth of the family Ypsolophidae first described by William D. Kearfott in 1907. It is found throughout North America. In Canada, it is known from British Columbia, Alberta and Saskatchewan. The wingspan T ...
'' (Kearfott, 1907) *'' Ypsolopha electropa'' (Meyrick, 1914) *'' Ypsolopha elongata'' (Braun, 1925) *'' Ypsolopha ephedrella'' (Christoph, 1873) *''
Ypsolopha excisella ''Ypsolopha excisella'' is a moth of the family Ypsolophidae. It is known from Spain, but has also been recorded from Mongolia and Asia Minor Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a ...
'' (Lederer, 1855) *'' Ypsolopha exsularis'' (Meyrick, 1937) *''
Ypsolopha falcella ''Ypsolopha falcella'' is a moth of the family Ypsolophidae. It is known from northern and central Europe and Russia. The wingspan is 17–20 mm. The antennas lies just over half the length of front wing. A wide, yellow-white longitudinal l ...
'' (Denis & Schiffermüller, 1775) *'' Ypsolopha falciferella'' (Walsingham, 1881) *''
Ypsolopha flavistrigella ''Ypsolopha flavistrigella'' is a moth of the family Ypsolophidae first described by August Busck in 1906. It was known only from the southern part of the United States (Texas, California, Utah and Colorado), but has also been recorded from Albe ...
'' (Busck, 1906) *'' Ypsolopha flava'' (Issiki, 1930) *'' Ypsolopha fractella'' (Chrétien, 1915) *''
Ypsolopha frustella ''Ypsolopha frustella'' is a moth of the family Ypsolophidae. It is known from the United States, including California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pac ...
'' (Walsingham, 1881) *''
Ypsolopha fujimotoi ''Ypsolopha fujimotoi'' is a moth of the family Ypsolophidae. It is known from Japan. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–20 ...
'' Moriuti, 1964 *'' Ypsolopha gerdanella'' (Busck, 1903) *'' Ypsolopha helva'' J.C. Sohn & C.S. Wu, in Sohn et al., 2010 *''
Ypsolopha heteraula ''Ypsolopha heteraula'' is a moth of the family Ypsolophidae. It is known from North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by th ...
'' (Meyrick, 1927) *'' Ypsolopha horridella'' (Tritschke, 1835) *''
Ypsolopha indecorella ''Ypsolopha indecorella'' is a moth of the family Ypsolophidae. It is known from France. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777 ...
'' (Rebel, 1903) *''
Ypsolopha instabilella ''Ypsolopha instabilella'' is a moth of the family Ypsolophidae. It is found in Uzbekistan, Tajikistan, Kyrghyzstan, Asia Minor and in the southern part of eastern Europe. The larvae feed on ''Ephedra'' species, including ''Ephedra distachya ...
'' (Mann, 1866) *'' Ypsolopha japonica'' Moriuti, 1964 *''
Ypsolopha kristalleniae ''Ypsolopha kristalleniae'' is a moth of the family Ypsolophidae. It is known from Crete and Turkey. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the ...
'' Rebel, 1916 *'' Ypsolopha leptaula'' (Meyrick, 1927) *'' Ypsolopha leuconotella'' (Snellen, 1884) *'' Ypsolopha lonicerella'' Stökl, 1922 *'' Ypsolopha longa'' Moriuti, 1964 *'' Ypsolopha lucella'' (Fabricius, 1775) *''
Ypsolopha lutisplendida ''Ypsolopha lutisplendida'' is a moth of the family Ypsolophidae. It is known from north-western China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a po ...
'' Sohn & Wu, 2011 *'' Ypsolopha lyonothamnae'' (Powell, 1967) *'' Ypsolopha maculatella'' (Busck, 1906) *''
Ypsolopha manella ''Ypsolopha manella'' is a moth of the family Ypsolophidae. It is known from the United States, including Utah and California California is a U.S. state, state in the Western United States, located along the West Coast of the United State ...
'' (Busck, 1903) *''
Ypsolopha manniella ''Ypsolopha manniella'' is a moth of the family Ypsolophidae. It is known from Crete and Turkey. The wingspan is 14–16 mm. The larvae feed on ''Ephedra Ephedra may refer to: * Ephedra (medicine), a medicinal preparation from the plant ' ...
'' (Staudinger, 1880) *''
Ypsolopha melanocnista ''Ypsolopha melanocnista'' is a moth of the family Ypsolophidae. It is known from Yunnan in China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a populat ...
'' (Meyrick, 1938) *''
Ypsolopha melanofuscella ''Ypsolopha melanofuscella'' is a moth of the family Ypsolophidae. It is found in the Russian Far East. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, th ...
'' Ponomarenko & Zinchenko, 2013 *''
Ypsolopha mienshani ''Ypsolopha mienshani'' is a moth of the family Ypsolophidae. It is known from Shanxi in China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population ...
'' (Caradja, 1939) *''
Ypsolopha minotaurella ''Ypsolopha minotaurella'' is a moth of the family Ypsolophidae. It is known from Albania, Greece and Crete. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For exampl ...
'' (Rebel, 1916) *''
Ypsolopha mucronella ''Ypsolopha mucronella'' is a moth of the family Ypsolophidae. It is found from Europe, through Siberia to Japan and in Asia Minor. The wingspan is 26–33 mm. The forewings are narrow, apex very strongly and acutely produced; light brownis ...
'' (Scopoli, 1763) *''
Ypsolopha nebulella ''Ypsolopha nebulella'' is a moth of the family Ypsolophidae. It is known from Russia. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–2 ...
'' (Staudinger, 1871) *''
Ypsolopha nella ''Ypsolopha nella'' is a moth of the family Ypsolophidae. It is known from the United States, including Arizona and Utah. The wingspan is about 21 mm. The antennae are white with narrow sharp black annulations. The labial palpi are dark ca ...
'' (Busck, 1903) *''
Ypsolopha nemorella ''Ypsolopha nemorella'' is a moth of the family Ypsolophidae. It is found in northern and central Europe, mid-eastern China and Russia. The wingspan is 21–24 mm. Adults are on wing from July to August. It is one of the larger species in t ...
'' (Linnaeus, 1758) *'' Ypsolopha nigrimaculata'' Byun et Park, 2001 *''
Ypsolopha nigrofasciata ''Ypsolopha nigrofasciata'' is a moth of the family Ypsolophidae. It is known from China (Hebei) and the Russian Far East The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐ ...
'' Yang, 1977 *''
Ypsolopha oliviella ''Ypsolopha oliviella'' is a moth of the family Ypsolophidae. It is known from the United States, including Arizona. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. Fo ...
'' (Busck, 1903) *''
Ypsolopha parallela ''Ypsolopha parallela'' is a moth of the family Ypsolophidae. It is known from Japan, Korea, China and Russia. The wingspan is 20–24 mm. The larvae feed on ''Quercus serrata ''Quercus serrata'', the jolcham oak, (, ) is an East Asian ...
'' (Caradja, 1939) *''
Ypsolopha parenthesella ''Ypsolopha parenthesella'' is a moth of the family Ypsolophidae. It is found from Europe to Japan, as well as north-eastern China, Asia Minor and mideast Asia. The wingspan is 16–20 mm. Adults are on wing from August to September. It is ...
'' (Linnaeus, 1761) *'' Ypsolopha parodaula'' (Meyrick, 1938) *''
Ypsolopha persicella ''Ypsolopha persicella'' is a moth of the family Ypsolophidae. It is found in southern-eastern and central Europe, the Crimea, the Caucasus and Asia Minor. The wingspan is 19–21 mm. The larvae feed on Rosaceae species, including almond, ...
'' (Fabricius, 1787) *'' Ypsolopha pseudoparallela'' J.C. Sohn & C.S. Wu, in Sohn et al., 2010 *'' Ypsolopha querciella'' (Busck, 1903) *''
Ypsolopha rubrella ''Ypsolopha rubrella'' is a moth of the family Ypsolophidae. It is found in the United States from California east to Colorado and Oklahoma. The wingspan is about 18 mm. The larvae feed on '' Berberis repens'' in Colorado and ''Berberis ne ...
'' (Dyar, 1902) *''
Ypsolopha rhytidota ''Ypsolopha rhytidota'' is a moth of the family Ypsolophidae. It is known from Yunnan in China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population ...
'' (Meyrick, 1938) *'' Ypsolopha saitoi'' Moriuti, 1964 *'' Ypsolopha sarmaticella'' (Rebel, 1917) *''
Ypsolopha sasayamanus ''Ypsolopha sasayamana'' is a moth of the family Ypsolophidae. It has been found in Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bo ...
'' (Matsumura, 1931) *''
Ypsolopha satellitella ''Ypsolopha satellitella'' is a moth of the family Ypsolophidae. It is known from Turkmenistan, Kyrghyzstan, north-western China, Afghanistan, the European part of Russia and Türkiye. The wingspan The wingspan (or just span) of a bird or ...
'' (Staudinger, 1871) *''
Ypsolopha scabrella ''Ypsolopha scabrella'', the wainscot hooktip or wainscot smudge, is a moth of the family Ypsolophidae. The species was first described by Carl Linnaeus in 1761. It is found in Europe, China, Russia, Asia Minor and mideast Asia. The wingspan ...
'' (Linnaeus, 1761) *'' Ypsolopha schwarziella'' (Busck, 1903) *''
Ypsolopha sculpturella ''Ypsolopha sculpturella'' is a moth of the family Ypsolophidae. It is known from Croatia, Bulgaria, Ukraine, Greece, Crete, Turkey and Israel. The larvae have been recorded feeding on ''Ephedra distachya ''Ephedra distachya'' is a shrub in t ...
'' (Herrich-Schäffer, 1854) *''
Ypsolopha semitessella ''Ypsolopha semitessella'' is a moth of the family Ypsolophidae. It is known from Croatia and North Macedonia North Macedonia, ; sq, Maqedonia e Veriut, (Macedonia before February 2019), officially the Republic of North Macedonia,, i ...
'' (Mann, 1861) *'' Ypsolopha senex'' (Walsingham, 1889) *''
Ypsolopha seniculella ''Ypsolopha seniculella'' is a moth of the family Ypsolophidae. It is known from Turkmenistan and Kyrghyzstan Kyrgyzstan,, pronounced or the Kyrgyz Republic, is a landlocked country in Central Asia. Kyrgyzstan is bordered by Kazakhstan ...
'' (Christoph, 1872) *''
Ypsolopha sequella ''Ypsolopha sequella'' is a moth of the family Ypsolophidae. It is found in Europe and Anatolia. The wingspan is 18–20 mm. The moth flies from July to September depending on the location. The larvae feed on willow, maple and ''Tilia ...
'' (Clerck, 1759) *''
Ypsolopha sordida ''Ypsolopha sordida'' is a moth of the family Ypsolophidae. It is known from southern China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population ex ...
'' J.C. Sohn & C.S. Wu, in Sohn et al., 2010 *''
Ypsolopha straminella ''Ypsolopha straminella'' is a moth of the family Ypsolophidae. It is found in the Russian Far East. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the ...
'' Ponomarenko & Zinchenko, 2013 *''
Ypsolopha striatella ''Ypsolopha striatella'' is a moth of the family Ypsolophidae. It is known from the United States, including California. The wingspan is about 22 mm. The antennae are white towards the apex with dark brown annulations. The labial palpi, es ...
'' (Busck, 1903) *''
Ypsolopha strigosus ''Ypsolopha strigosa'' is a moth of the family Ypsolophidae. It is known from Japan, Korea, China and Russia. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For examp ...
'' (Butler, 1879) *''
Ypsolopha sublucella ''Ypsolopha sublucella'' is a moth of the family Ypsolophidae. It is known from California, United States. The larvae feed on oaks of the section '' Protobalanus'' (intermediate oaks), including ''Quercus vaccinifolia ''Quercus vacciniifolia' ...
'' (Walsingham, 1881) *''
Ypsolopha sylvella ''Ypsolopha sylvella'' is a moth of the family Ypsolophidae. It is found in most of Europe. The wingspan is 18–20 mm. Adults are on wing from August to September. There is one generation per year. The larvae feed on the leaves of ''Querc ...
'' (Linnaeus, 1767) *''
Ypsolopha tesselatidorsata ''Ypsolopha tesselatidorsata'' is a moth of the family Ypsolophidae. It is known from the Russian Far East The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲi ...
'' Ponomarenko & Zinchenko, 2011 *'' Ypsolopha trichonella'' (Mann, 1861) *'' Ypsolopha tsugae'' Moriuti, 1977 *'' Ypsolopha undulatella'' (Busck, 1906) *''
Ypsolopha unicipunctella ''Ypsolopha unicipunctella'' is a moth of the family Ypsolophidae. It is known from the United States, including Oklahoma, Texas and Arizona. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip ...
'' (Busck, 1903) *''
Ypsolopha uniformis ''Ypsolopha uniformis'' is a moth of the family Ypsolophidae. It was described by Ivan Nikolayevich Filipjev in 1929 and is known from Buryatia, a republic of Russia Russia (, , ), or the Russian Federation, is a List of transcontinenta ...
'' (Filipjev, 1929) *''
Ypsolopha ustella ''Ypsolopha ustella'', the variable ypsolopha moth, is a moth of the family Ypsolophidae. It is found in most of Europe and is also present in North America. The wingspan is 15–20 mm. It is a variable species with numerous colour forms ...
'' (Clerck, 1759) *''
Ypsolopha vintrella ''Ypsolopha vintrella'' is a moth of the family Ypsolophidae. It is known from the United States, including Arizona and California. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the ot ...
'' (Busck, 1906) *''
Ypsolopha vittella ''Ypsolopha vittella'', the elm autumn moth, is a moth of the family Ypsolophidae. It is found from Europe through Siberia to Japan, including China, Asia Minor and mideast Asia. The habitat consists of woodlands and copses. The wingspan is 16 ...
'' (Linnaeus, 1758) *'' Ypsolopha walsinghamiella'' (Busck, 1903) *''
Ypsolopha yangi ''Ypsolopha yangi'' is a moth of the family Ypsolophidae. It is known from the Russian Far East and northern and south-western China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's mo ...
'' Ponomerenko & Sohn, 2011 *'' Ypsolopha yasudai'' Moriuti, 1964


Former species

*'' Ypsolopha scenites'' (Meyrick, 1909)


References

{{Taxonbar, from=Q3020455 Ypsolophidae Moth genera Taxa named by Pierre André Latreille