The yellow longnose butterflyfish or forceps butterflyfish (''Forcipiger flavissimus'') is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of marine

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Chaetodontidae. The yellow longnose butterflyfish is widespread throughout the

tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...

waters of the

Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...

area from the eastern coast of

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...

Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; T ...

included, and is also found in the eastern

Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...

from

Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...

to the

Revillagigedo Islands The Revillagigedo Islands ( es, Islas Revillagigedo, ) or Revillagigedo Archipelago are a group of four volcanic islands in the Pacific Ocean, known for their unique ecosystem. They lie approximately from Socorro Island south and southwest of C ...

and the Galapagos. It is a small

which grows up to in length.Lieske & Myers,''Coral reef fishes'',Princeton University Press, 2009, This species can also be found in the

aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...

trade.

Territoriality

Being

territorial A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...

, yellow longnose butterflyfish patrol their patches of coral with a

monogamous Monogamy ( ) is a form of Dyad (sociology), dyadic Intimate relationship, relationship in which an individual has only one Significant other, partner during their lifetime. Alternately, only one partner at any one time (Monogamy#Serial monogamy, ...

partner. However, instances of overt

aggression Aggression is overt or covert, often harmful, social interaction with the intention of inflicting damage or other harm upon another individual; although it can be channeled into creative and practical outlets for some. It may occur either reacti ...

among ''F. flavissimus'' have been observed between territory holders and individuals of the same sex. Chasing is rare, but when it does occur, males chase males and females chase females. Females defend food resources from other females, while males defend territories containing a female from other males.

Territoriality In ethology, territory is the sociographical area that an animal consistently defends against conspecific competition (or, occasionally, against animals of other species) using agonistic behaviors or (less commonly) real physical aggression. ...

is a favorable strategy for a species to adopt primarily when resources are temporally stable, predictable, and evenly distributed throughout a territory. Territoriality is commonly displayed by benthic-feeding longnose butterflyfish, therefore, because their main dietary resources fulfill these characteristics. Their monogamous pairing appears to be closely linked to their territorial behavior. Although several could cause a species to evolve monogamous behavior, the necessity for biparental care does not apply to longnose butterflyfish because they lay pelagic, or freely floating, eggs. One source of selective pressure responsible for the monogamous pairs observed could be the advantage of territorial defense it provides. Monogamy is favored when a pair makes the defense of one or more resources more efficient than defense by a solitary individual. Longnose butterflyfish pairs have been confirmed by studies to be

heterosexual Heterosexuality is romantic attraction, sexual attraction or sexual behavior between people of the opposite sex or gender. As a sexual orientation, heterosexuality is "an enduring pattern of emotional, romantic, and/or sexual attractions" to ...

and pair

fidelity Fidelity is the quality of faithfulness or loyalty. Its original meaning regarded duty in a broader sense than the related concept of ''fealty''. Both derive from the Latin word ''fidēlis'', meaning "faithful or loyal". In the City of London f ...

has been observed for periods of up to seven or more years. Besides the advertisement displays accomplished through monogamous pairing, territorial domination by longnose butterflyfish has also been observed by means of acoustic behaviors, which provide important cues and social signals during fish communication. Emitting sounds through complicated body movements is another technique they use to advertise territorial boundaries. Potential rivals are able to assess body size of a competitor based on the duration and intensity of the sound a yellow longnose butterflyfish produces. The duration and intensity of the sounds emitted during antagonistic behaviors, such as the defense of one's territory, often predict the ability of an individual to secure that territory. A sound of long duration and high intensity, therefore, often indicates an individual has a large territory. Defending territory is the strategy these species adopt to compete for and maximize their claims over resources.

References

External links

*
WoRMS - World Register of Marine Species - Forcipiger flavissimus Jordan & McGregor, 1898
* * Photos o
Forcipiger flavissimus
in iNaturalist {{Taxonbar, from=Q1499346 Forcipiger Fish of the Pacific Ocean Fish of Central America Fish of Hawaii Fish of Mexican Pacific coast Galápagos Islands coastal fauna Natural history of the Revillagigedo Islands Fish described in 1898 Taxa named by David Starr Jordan