Yellow-legged Dragonfly
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The river clubtail or yellow-legged dragonfly (''Gomphus flavipes'') is a species of
dragonfly A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of true dragonfly are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threate ...
in the family Gomphidae. It is found in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
. Its natural
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
are rivers and large streams. The dragonfly flies from June to September depending on the location.


Geographical distribution

''G. flavipes'' is a European species. It has the biggest geographical range among the European Gomphid species, occurring from
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
to eastern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
. The species has a very patchy occurrence in Central Europe and has become rare in Western Europe. The southernmost occurrence is known from Greece. In Southern Europe ''G. flavipes'' is replaced by other Gomphid species. However, the exact geographical range of the species is still unclear.


Habitat

''G. flavipes'' lives along the middle and lower reaches of slow flowing medium-sized and big rivers The species prefers riverbed with mud, clay, loam or fine sand as soil, since larvae develop buried in the soil, but avoids areas with decomposition of organic matter and oxygen-poor
stagnant water Water stagnation occurs when water stops flowing. Stagnant water can be a major environmental hazard. Dangers Malaria and dengue are among the main dangers of stagnant water, which can become a breeding ground for the mosquitoes that transmi ...
. Reaches with high drift and coarse substrate are also avoided, however, occasionally ''G. flavipes'' can be found in bigger lakes with high oxygen content.


Morphology

As a medium-sized dragonfly ''G. flavipes'' is 50–55 mm long and with an average wingspan of 70–80 mm. The eyes are widely separated, this is a typical genus-specific feature, males have blue while females have green eyes. Both sexes have predominantly yellow legs and the males has thin angled appendages, while his 10th segment is entirely black dorsally.


Behaviour and life cycle

''G. flavipes'', as all dragonflies, are predators; they capture smaller insect in flight.
Imago In biology, the imago (Latin for "image") is the last stage an insect attains during its metamorphosis, its process of growth and development; it is also called the imaginal stage, the stage in which the insect attains maturity. It follows the f ...
s are rarely seen, the easiest way to observe them is in the mating period or during the emergence of young adults. Mating usually takes place in June or July. During the day adult males search for females, far from the riverbank, in low altitude above the waterfront. If they successfully catch a female copulation takes place in the shelter of water edge willows. After fertilization females fly 20–30 cm above the water and lay eggs (one at a time or in a small packets) into the water.


Larval development

The
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e develop for three years buried in the fine soil of the riverbed. As imagos, they are predators too, their diet consisting of small invertebrates which they capture with they specialized labium called the ''mask''. The emergence of the larvae starts mostly at end of May or June, but it depends highly on location and other environmental circumstances (e.g. a rainy period could delay the start of emergence). The emergence in ''G. flavipes'' is often highly synchronised and it is therefore referred as a ’summer species’, in Odonatology EM50 values are used to describe the emergence pattern (extent of synchronization). The patterns often have two peaks in ''G. flavipes'' due to the phenomenon called ’cohort splitting’. Larvae of ''G. flavipes'', unlike other Anisoptera larvae, often choose riverbank soil as an emergence substrate, however the type of substrate depends on location and characteristics of the riverbank, probably the larvae have no substrate specific attachment.


Threat factors

''G. flavipes'' has become an endangered species in most Western European countries due to water pollution and river regulation. The species is listed in the Annex IV of Habitats Directive.


References


External links

* {{Taxonbar, from=Q728093 Gomphidae Dragonflies of Europe Odonata of Asia Insects described in 1825 Taxa named by Toussaint de Charpentier