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The yellow-billed loon (''Gavia adamsii''), also known as the white-billed diver, is the largest member of the
loon Loons (North American English) or divers ( British / Irish English) are a group of aquatic birds found in much of North America and northern Eurasia. All living species of loons are members of the genus ''Gavia'', family Gaviidae and order G ...
or diver family. Breeding adults have a black head, white underparts and chequered black-and-white mantle. Non-breeding
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
is drabber with the chin and foreneck white. Its main distinguishing feature is the long straw-yellow bill which, because the culmen is straight, appears slightly uptilted. It breeds in the
Arctic The Arctic ( or ) is a polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada ( Yukon, Northwest Territories, Nunavut), Danish Realm ( Greenland), Finland, Iceland ...
and winters mainly at sea along the
coast The coast, also known as the coastline or seashore, is defined as the area where land meets the ocean, or as a line that forms the boundary between the land and the coastline. The Earth has around of coastline. Coasts are important zones in n ...
s of the northern
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contine ...
and northwestern
Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and t ...
; it also sometimes overwinters on large inland lakes. It occasionally strays well south of its normal wintering range, and has been recorded as a vagrant in more than 22 countries. This
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
, like all divers, is a specialist
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% ...
-eater, catching its
prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
underwater. Its call is an eerie wailing, lower pitched than the common loon.


Taxonomy and etymology

First described by English zoologist
George Robert Gray George Robert Gray FRS (8 July 1808 – 6 May 1872) was an English zoologist and author, and head of the ornithological section of the British Museum, now the Natural History Museum, in London for forty-one years. He was the younger broth ...
in 1859 based on a specimen collected in Alaska, the yellow-billed loon is a monotypic species, with no subspecies despite its large
Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical regi ...
range. It is closely related to the
common loon The common loon or great northern diver (''Gavia immer'') is a large member of the loon, or diver, family of birds. Breeding adults have a plumage that includes a broad black head and neck with a greenish, purplish, or bluish sheen, blackish ...
, which it strongly resembles in plumage and behaviour; some taxonomists consider the two species to be
allopatric Allopatric speciation () – also referred to as geographic speciation, vicariant speciation, or its earlier name the dumbbell model – is a mode of speciation that occurs when biological populations become geographically isolated from ...
forms of the same superspecies. Both are thought to have evolved from a population of black-throated loons which colonized the
Nearctic The Nearctic realm is one of the eight biogeographic realms constituting the Earth's land surface. The Nearctic realm covers most of North America, including Greenland, Central Florida, and the highlands of Mexico. The parts of North America ...
and were subsequently cut off from other populations. The genus name ''Gavia'' comes from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
for "sea mew", as used by ancient Roman naturalist
Pliny the Elder Gaius Plinius Secundus (AD 23/2479), called Pliny the Elder (), was a Roman author, naturalist and natural philosopher, and naval and army commander of the early Roman Empire, and a friend of the emperor Vespasian. He wrote the encyclopedic ...
. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''adamsii'' honours Edward Adams, a British
naval surgeon A naval surgeon, or less commonly ship's doctor, is the person responsible for the health of the ship's company aboard a warship. The term appears often in reference to Royal Navy's medical personnel during the Age of Sail. Ancient uses Speciali ...
and naturalist who sketched and collected numerous species, including this one, on several trips to the
Arctic The Arctic ( or ) is a polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada ( Yukon, Northwest Territories, Nunavut), Danish Realm ( Greenland), Finland, Iceland ...
. The word "loon" is thought to have derived from the Swedish ''lom'', the Old Norse or Icelandic ''lómr'', or the Old Dutch ''loen'', all of which mean "lame" or "clumsy", and is a probable reference to the difficulty that all loons have in moving about on land. "Diver" refers to the family's underwater method of hunting for prey, while "yellow-billed" and "white-billed" are references to the bird's distinctively pale bill.


Description

With a length of , a wingspan of , and a weight ranging from , the yellow-billed loon is the largest member of the loon (diver) family. The adult is primarily black and white in breeding
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
, with a purple gloss on its head and neck.


Habitat and range

The yellow-billed loon is an Arctic species, breeding primarily along the coasts of the Arctic Ocean as far north as 78° N and wintering on sheltered coastal waters of the northern
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contine ...
and the northwestern coast of
Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and t ...
. It has been recorded as a breeding bird in
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ...
, Canada and the United States. Though it winters primarily to the north of 50° N, its winter range extends south to 35° N off the coast of Japan, and it has been recorded as a vagrant in more than 20 countries, including some as far south as
Mexico Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...
and
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' ( Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
.


Behaviour


Breeding

Like other loons, it forms long-lasting pairs. Though it prefers freshwater pools or lakes in the tundra, the yellow-billed loon will also breed along rivers, estuaries or the coast in low-lying areas of the Arctic; in general, it avoids forested areas. Breeding typically starts in early June, though it is dependent on the timing of the spring thaw. Like all members of its family, the yellow-billed loon builds a
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materi ...
of plant material very close to the edge of the water. Copulation takes place on land, without any specific courtship. The pair defends its large territory intensively against intruders, but may later in the breeding season gang up with other birds on good fishing spots. The female lays two eggs measuring . The eggs are strongly oval, and are a light purple-brown with darker blotches interspersed. This colour camouflages with the soil and vegetation that this bird nests near. Most of the time, however, the egg is not visible due to incubation, which takes around 27 to 29 days.


Feeding

The yellow-billed loon is a specialist fish eater, yet it also takes
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean gro ...
s, molluscs and
annelid The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecol ...
s, especially for its young. It dives in pursuit of prey, which is caught underwater. Probably as a way to avoid spreading parasites, it defecates ashore, in the breeding lake.


Conservation and threats

In 2010, the IUCN (International Union for the Conservation of Nature) changed the status of the yellow-billed loon from
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. Th ...
to
Near Threatened A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify fo ...
, as the species appears to be in a "moderately rapid" population decline. An unsustainable level of subsistence harvesting by
indigenous people Indigenous peoples are culturally distinct ethnic groups whose members are directly descended from the earliest known inhabitants of a particular geographic region and, to some extent, maintain the language and culture of those original people ...
s was specifically named as the main threat. The yellow-billed loon is one of the
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
to which the ''
Agreement on the Conservation of African-Eurasian Migratory Waterbirds The Agreement on the Conservation of African-Eurasian Migratory Waterbirds, or African-Eurasian Waterbird Agreement (AEWA) is an independent international treaty developed under the auspices of the United Nations Environment Programme's Convent ...
'' (AEWA) applies; in the Americas, it is protected by the
Migratory Bird Treaty Act of 1918 The Migratory Bird Treaty Act of 1918 (MBTA), codified at (although §709 is omitted), is a United States federal law, first enacted in 1918 to implement the convention for the protection of migratory birds between the United States and Canada . ...
.


References


Citations


Bibliography

Sjölander, Sverre. The reproductive Behavior of the Yellow-billed Loon, Gavia adamsii. (With G. Ågren). The Condor 78:454-463. 1976.


Further reading

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Identification

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External links


Flicker Field Guide Birds of the World
Photographs * * * * * {{Authority control yellow-billed loon Birds of the Arctic Holarctic birds yellow-billed loon yellow-billed loon