Xenus Cinereus
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The Terek sandpiper (''Xenus cinereus'') is a small migratory
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
wader species and is the only member of the genus ''Xenus''. It is named after the Terek River which flows into the west of the Caspian Sea, as it was first observed around this area.


Taxonomy

The Terek sandpiper was formally described and illustrated in 1775 by the German naturalist Johann Anton Güldenstädt under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Scolopax cinerea''. He reported that he had seen pairs breeding at the mouth of the Terek River where it flows into the Caspian Sea. It is now the only species placed in the genus ''Xenus'' that was introduced in 1829 by the German naturalist
Johann Jakob Kaup Johann Jakob von Kaup (10 April 1803 – 4 July 1873) was a German naturalist. A proponent of natural philosophy, he believed in an innate mathematical order in nature and he attempted biological classifications based on the Quinarian system. Kaup ...
. The genus name ''Xenus'' is from Ancient Greek ''xenos'' meaning "stranger"; the specific epithet ''cinereus'' is Latin for "ash-grey" from ''cinis, cineris'', "ashes". The species is considered to be monotypic, no
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised. Among the
Scolopacidae Sandpipers are a large family, Scolopacidae, of waders. They include many species called sandpipers, as well as those called by names such as curlew and snipe. The majority of these species eat small invertebrates picked out of the mud or soil. ...
, ''Xenus'' is part of the shank- tattler-
phalarope __NOTOC__ A phalarope is any of three living species of slender-necked shorebirds in the genus ''Phalaropus'' of the bird family Scolopacidae. Phalaropes are close relatives of the shanks and tattlers, the ''Actitis'' and Terek sandpipers, a ...
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
and less closely related to the calidrid sandpipers. Based on the degree of
DNA sequence DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, guanine, cytosine, and thymine. Th ...
divergence and putative shank and phalarope fossils from around the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
/ Miocene boundary some 23–22 million years ago, the Terek sandpiper presumably diverged from their relatives in the Late Oligocene. Given the numerous
basal Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''. Science * Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure * Basal (medicine), a minimal level that is nec ...
fossils of the group found in Eurasia it is likely that the Terek sandpiper lineage originated there, possibly by being isolated as the remains of the Turgai Sea dried up, which happened just around this time.


Description

Slightly larger than the
common sandpiper The common sandpiper (''Actitis hypoleucos'') is a small Palearctic wader. This bird and its American sister species, the spotted sandpiper (''A. macularia''), make up the genus ''Actitis''. They are parapatric and replace each other geographic ...
at length, its long upcurved bill – somewhat reminiscent of an avocet's, but not as strongly curved – makes it very distinctive. As the scientific specific name implies, this wader has a grey back, face and breast in all
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
s; a white
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also ...
may appear more or less distinct. The belly is whitish and the feet yellow; the
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
has a yellowish base, with the rest being black. The call is a high whistle.


Distribution and ecology

This bird breeds near water in the taiga from Finland through northern Siberia to the Kolyma River, and
migrate Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...
south in winter to tropical coasts in east Africa, south Asia and
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, usually preferring muddy areas. It is a rare vagrant in western Europe, and particularly in autumn it is sometimes seen passing through the
Marianas The Mariana Islands (; also the Marianas; in Chamorro: ''Manislan Mariånas'') are a crescent-shaped archipelago comprising the summits of fifteen longitudinally oriented, mostly dormant volcanic mountains in the northwestern Pacific Ocean, betw ...
on migration; on Palau, further off its usual migration route, it is decidedly uncommon on the other hand. Almost annually and apparently more and more often in recent times, a few birds stray to Alaska and the Aleutian and Pribilof Islands. Every few years, individual vagrants are recorded in the Neotropics, where they arrive either as migrating birds from Africa, or as North American strays accompanying local waders south for winter. Such vagrants have been recorded as far south as Argentina. The overall genetic variation in Terek sandpipers across their range is low, with some evidence of contractions followed by expansion. Although the geographically isolated Dniephe River population in Eastern Europe does show significant genetic differentiation. It feeds in a distinctive and very active way, chasing insects and other mobile prey, and sometimes then running to the water's edge to wash its catch. It lays three or four eggs in a lined ground scrape. The Terek sandpiper likes to associate with ruddy turnstones (''Arenaria interpres''), smallish calidrids, and '' Charadrius'' (but maybe not '' Pluvialis'') plovers; a vagrant bird at Paraty ( Rio de Janeiro state) was noted to pair up with a
spotted sandpiper The spotted sandpiper (''Actitis macularius'') is a small shorebird. Together with its sister species the common sandpiper (''A. hypoleucos''), it makes up the genus ''Actitis''. They replace each other geographically; stray birds may settle dow ...
(''Actitis macularius''). This is one of the species to which the ''Agreement on the Conservation of African-Eurasian Migratory Waterbirds'' ( AEWA) applies. Widespread and often quite commonly seen, the Terek sandpiper is not considered a threatened species by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
.


Gallery

File:Terek Sandpiper, Little Stint, Marsh Sandpiper, C Redshank W2 IMG 6891.jpg, Terek sandpiper ''Xenus cinereus'', little stint ''Calidris minuta'' and
marsh sandpiper The marsh sandpiper (''Tringa stagnatilis'') is a small wader. It is a rather small shank, and breeds in open grassy steppe and taiga wetlands from easternmost Europe to the Russian Far East. The genus name ''Tringa'' is the New Latin name given ...
''Tringa stagnatilis'' in Krishna Wildlife Sanctuary, Andhra Pradesh, India. File:Xenus cinereus eating ragworm.JPG, Eating ragworm in Shōnai River,
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
. File:Xenus cinereus MWNH 0248.JPG, Egg, Collection Museum Wiesbaden File:Terek Sandpiper (Xenus cinereus) (8079446567).jpg, In flight File:Terek Sandpiper AMSM5480 TSAN.jpg, In Narara,
Jamnagar Jamnagar () is a city located on the western coast of India in the state of Gujarat of Saurashtra (region), Saurashtra region. It is the administrative headquarters of the Jamnagar district and the fifth largest city in Gujarat. The city lies ...
, India File:Terek Sandpiper AMSM5481.jpg, In Narara,
Jamnagar Jamnagar () is a city located on the western coast of India in the state of Gujarat of Saurashtra (region), Saurashtra region. It is the administrative headquarters of the Jamnagar district and the fifth largest city in Gujarat. The city lies ...
, India


References


External links


Terek sandpiper species text in The Atlas of Southern African Birds
* * * * * * * {{Taxonbar, from=Q18856 Terek sandpiper Birds of Russia Birds of Central Asia Terek sandpiper Terek sandpiper