Xenus (bird)
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The Terek sandpiper (''Xenus cinereus'') is a small migratory
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
wader 245px, A flock of Dunlins and Red knots">Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflat ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
and is the only member of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Xenus''. It is named after the Terek River which flows into the west of the
Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad steppe of Central Asia ...
, as it was first observed around this area.


Taxonomy

The Terek sandpiper was formally described and illustrated in 1775 by the German naturalist Johann Anton Güldenstädt under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Scolopax cinerea''. He reported that he had seen pairs breeding at the mouth of the Terek River where it flows into the
Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad steppe of Central Asia ...
. It is now the only species placed in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Xenus'' that was introduced in 1829 by the German naturalist
Johann Jakob Kaup Johann Jakob von Kaup (10 April 1803 – 4 July 1873) was a German naturalist. A proponent of natural philosophy, he believed in an innate mathematical order in nature and he attempted biological classifications based on the Quinarian system. Kaup ...
. The genus name ''Xenus'' is from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''xenos'' meaning "stranger"; the specific epithet ''cinereus'' is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for "ash-grey" from ''cinis, cineris'', "ashes". The species is considered to be
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
, no
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised. Among the
Scolopacidae Sandpipers are a large family, Scolopacidae, of waders. They include many species called sandpipers, as well as those called by names such as curlew and snipe. The majority of these species eat small invertebrates picked out of the mud or soil. ...
, ''Xenus'' is part of the shank- tattler-
phalarope __NOTOC__ A phalarope is any of three living species of slender-necked shorebirds in the genus ''Phalaropus'' of the bird family Scolopacidae. Phalaropes are close relatives of the shanks and tattlers, the ''Actitis'' and Terek sandpipers, a ...
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
and less closely related to the calidrid sandpipers. Based on the degree of
DNA sequence DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, guanine, cytosine, and thymine. Th ...
divergence and putative shank and phalarope
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s from around the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
/
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
boundary some 23–22
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
, the Terek sandpiper presumably diverged from their relatives in the Late Oligocene. Given the numerous
basal Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''. Science * Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure * Basal (medicine), a minimal level that is nec ...
fossils of the group found in
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
it is likely that the Terek sandpiper lineage originated there, possibly by being isolated as the remains of the
Turgai Sea The Turgai Sea, also known as the Turgay Sea, Turgai Strait, Obik Sea, Ural Sea or West Siberian Sea, was a large shallow body of salt water (an epicontinental or epeiric sea) during the Mesozoic through Cenozoic Eras. It extended north of the pres ...
dried up, which happened just around this time.


Description

Slightly larger than the
common sandpiper The common sandpiper (''Actitis hypoleucos'') is a small Palearctic wader. This bird and its American sister species, the spotted sandpiper (''A. macularia''), make up the genus ''Actitis''. They are parapatric and replace each other geographic ...
at length, its long upcurved bill – somewhat reminiscent of an avocet's, but not as strongly curved – makes it very distinctive. As the scientific specific name implies, this wader has a grey back, face and breast in all
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
s; a white
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also ...
may appear more or less distinct. The belly is whitish and the feet yellow; the
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
has a yellowish base, with the rest being black. The call is a high whistle.


Distribution and ecology

This bird breeds near water in the
taiga Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruce ...
from
Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of B ...
through northern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
to the
Kolyma River The Kolyma ( rus, Колыма, p=kəlɨˈma; sah, Халыма, translit=Khalyma) is a river in northeastern Siberia, whose basin covers parts of the Sakha Republic, Chukotka Autonomous Okrug, and Magadan Oblast of Russia. The Kolyma is frozen ...
, and
migrate Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...
south in winter to tropical coasts in east Africa, south Asia and
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, usually preferring muddy areas. It is a rare vagrant in western Europe, and particularly in autumn it is sometimes seen passing through the
Marianas The Mariana Islands (; also the Marianas; in Chamorro: ''Manislan Mariånas'') are a crescent-shaped archipelago comprising the summits of fifteen longitudinally oriented, mostly dormant volcanic mountains in the northwestern Pacific Ocean, betw ...
on migration; on
Palau Palau,, officially the Republic of Palau and historically ''Belau'', ''Palaos'' or ''Pelew'', is an island country and microstate in the western Pacific. The nation has approximately 340 islands and connects the western chain of the Caro ...
, further off its usual migration route, it is decidedly uncommon on the other hand. Almost annually and apparently more and more often in recent times, a few birds stray to
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
and the Aleutian and
Pribilof Islands The Pribilof Islands (formerly the Northern Fur Seal Islands; ale, Amiq, russian: Острова Прибылова, Ostrova Pribylova) are a group of four volcanic islands off the coast of mainland Alaska, in the Bering Sea, about north of ...
. Every few years, individual vagrants are recorded in the Neotropics, where they arrive either as migrating birds from Africa, or as North American strays accompanying local waders south for winter. Such vagrants have been recorded as far south as
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. The overall genetic variation in Terek sandpipers across their range is low, with some evidence of contractions followed by expansion. Although the geographically isolated Dniephe River population in Eastern Europe does show significant genetic differentiation. It feeds in a distinctive and very active way, chasing insects and other mobile prey, and sometimes then running to the water's edge to wash its catch. It lays three or four eggs in a lined ground scrape. The Terek sandpiper likes to associate with ruddy turnstones (''Arenaria interpres''), smallish calidrids, and '' Charadrius'' (but maybe not ''
Pluvialis '' Pluvialis '' is a genus of plovers, a group of wading birds comprising four species that breed in the temperate or Arctic Northern Hemisphere. In breeding plumage, they all have largely black underparts, and golden or silvery upperparts. The ...
'')
plover Plovers ( , ) are a widely distributed group of wading birds belonging to the subfamily Charadriinae. Description There are about 66 species in the subfamily, most of them called "plover" or "dotterel". The closely related lapwing subfa ...
s; a vagrant bird at
Paraty Paraty (or Parati, ) is a preserved Portuguese colonial (1500–1822) and Brazilian Imperial (1822–1889) municipality with a population of about 43,000. Indeed, the name "Paraty" originates from the local Guaianá Indians' Indigenous Tupi l ...
(
Rio de Janeiro state Rio de Janeiro () is one of the 27 federative units of Brazil. It has the second largest economy of Brazil, with the largest being that of the state of São Paulo. The state, which has 8.2% of the Brazilian population, is responsible for 9.2% of ...
) was noted to pair up with a
spotted sandpiper The spotted sandpiper (''Actitis macularius'') is a small shorebird. Together with its sister species the common sandpiper (''A. hypoleucos''), it makes up the genus ''Actitis''. They replace each other geographically; stray birds may settle dow ...
(''Actitis macularius''). This is one of the species to which the ''Agreement on the Conservation of African-Eurasian Migratory Waterbirds'' ( AEWA) applies. Widespread and often quite commonly seen, the Terek sandpiper is not considered a threatened species by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
.


Gallery

File:Terek Sandpiper, Little Stint, Marsh Sandpiper, C Redshank W2 IMG 6891.jpg, Terek sandpiper ''Xenus cinereus'', little stint ''Calidris minuta'' and
marsh sandpiper The marsh sandpiper (''Tringa stagnatilis'') is a small wader. It is a rather small shank, and breeds in open grassy steppe and taiga wetlands from easternmost Europe to the Russian Far East. The genus name ''Tringa'' is the New Latin name given ...
''Tringa stagnatilis'' in
Krishna Wildlife Sanctuary Krishna Wildlife Sanctuary is a wildlife sanctuary and estuary located in Andhra Pradesh, India. It is one of the rarest eco-regions of the world because it harbors vast tracts of pristine mangrove forests. It is believed by conservationists to ...
,
Andhra Pradesh Andhra Pradesh (, abbr. AP) is a state in the south-eastern coastal region of India. It is the seventh-largest state by area covering an area of and tenth-most populous state with 49,386,799 inhabitants. It is bordered by Telangana to the ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
. File:Xenus cinereus eating ragworm.JPG, Eating ragworm in Shōnai River,
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
. File:Xenus cinereus MWNH 0248.JPG, Egg, Collection Museum Wiesbaden File:Terek Sandpiper (Xenus cinereus) (8079446567).jpg, In flight File:Terek Sandpiper AMSM5480 TSAN.jpg, In Narara,
Jamnagar Jamnagar () is a city located on the western coast of India in the state of Gujarat of Saurashtra (region), Saurashtra region. It is the administrative headquarters of the Jamnagar district and the fifth largest city in Gujarat. The city lies ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
File:Terek Sandpiper AMSM5481.jpg, In Narara,
Jamnagar Jamnagar () is a city located on the western coast of India in the state of Gujarat of Saurashtra (region), Saurashtra region. It is the administrative headquarters of the Jamnagar district and the fifth largest city in Gujarat. The city lies ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...


References


External links


Terek sandpiper species text in The Atlas of Southern African Birds
* * * * * * * {{Taxonbar, from=Q18856 Terek sandpiper Birds of Russia Birds of Central Asia Terek sandpiper Terek sandpiper