Xenotosuchus Skull Roof
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''Xenotosuchus'' is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of mastodonsaurid
temnospondyl Temnospondyli (from Greek language, Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order (biology), order of small to giant tetrapods—often considered Labyrinthodontia, primitive amphi ...
within the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Mastodonsauridae Mastodonsauridae is a family of capitosauroid temnospondyls. Fossils belonging to this family have been found in North America, Greenland, Europe, Asia, and Australia. The family Capitosauridae is synonymous with Mastodonsauridae. Descripti ...
known from the
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
of
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
. The genus is based on a skull originally described as ''
Parotosuchus ''Parotosuchus'' is an extinct genus of capitosaurian temnospondyl amphibians within the family Mastodonsauridae. Fossils are known from the Early Triassic of Europe, Africa, Australia, and Antarctica. It was about long and likely lived in aqua ...
'', an animal which it resembled in general build and habit.


Description

Like many mastodontosaurids, it was a large animal with a large head. Its
amphibian Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terres ...
life history meant that the distinct shape of the
skull roof The skull roof, or the roofing bones of the skull, are a set of bones covering the brain, eyes and nostrils in bony fishes and all land-living vertebrates. The bones are derived from dermal bone and are part of the dermatocranium. In comparati ...
would change from a generalized
tadpole A tadpole is the larval stage in the biological life cycle of an amphibian. Most tadpoles are fully aquatic, though some species of amphibians have tadpoles that are terrestrial. Tadpoles have some fish-like features that may not be found i ...
-like skull type through to the distinct adult shape. The head bones are covered in large pits and grooves, indicating extensive dermal armour on the head. Both the upper and lower jaw had
tusk Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine teeth, as with pigs and walruses, or, in the case of elephants, elongated incisors. Tusks share c ...
s, those of the upper jaw being situated on a 2nd row of teeth on the
vomer The vomer (; lat, vomer, lit=ploughshare) is one of the unpaired facial bones of the skull. It is located in the midsagittal line, and articulates with the sphenoid, the ethmoid, the left and right palatine bones, and the left and right maxill ...
and
palatine bone In anatomy, the palatine bones () are two irregular bones of the facial skeleton in many animal species, located above the uvula in the throat. Together with the maxillae, they comprise the hard palate. (''Palate'' is derived from the Latin ''pa ...
. Contrary to related forms like ''
Mastodonsaurus ''Mastodonsaurus'' (meaning "teat tooth lizard") is an extinct genus of temnospondyl amphibian from the Middle Triassic of Europe. It belongs to a Triassic group of temnospondyls called Capitosauria, characterized by their large body size and pre ...
'', the tusks of the lower jaw were of moderate size and did not penetrate the
premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has b ...
.


See also

*
Prehistoric amphibian This list of prehistoric amphibians is an attempt to create a comprehensive listing of all Genus, genera from the fossil record that have ever been considered to be amphibians, excluding purely vernacular terms. The list includes all commonly accep ...
*
List of prehistoric amphibians This list of prehistoric amphibians is an attempt to create a comprehensive listing of all Genus, genera from the fossil record that have ever been considered to be amphibians, excluding purely vernacular terms. The list includes all commonly accep ...


References

Early Triassic amphibians of Africa Triassic temnospondyls of Africa Fossil taxa described in 2002 Prehistoric amphibian genera {{Triassic-animal-stub