Woodhouse's toad

Woodhouse's toad (''Anaxyrus woodhousii'') is a medium-sized () true toad native to the United States and Mexico. There are three recognized subspecies. ''A. woodhousii'' tends to hybridize with '' Anaxyrus americanus'' where their ranges overlap.

Taxonomy

Woodhouse's toad was first described in 1854 by the French herpetologist Charles Frédéric Girard. He gave it the name ''Bufo woodhousii'' in honor of the American physician and naturalist Samuel Washington Woodhouse. The large genus ''Bufo'' was split by Frost ''et al.'' in 2006, with the North American species being included in the genus ''Anaxyrus'' and this toad becoming ''A. woodhousii''. There are three recognised subspecies:
*Rocky Mountain toad – '' Anaxyrus woodhousii woodhousii'' ( Girard, 1854)
*East Texas toad – ''Anaxyrus woodhousii velatus'' (Bragg and Sanders, 1951)Bragg, A. N., and O. Sanders. 1951. A new subspecies of the ''Bufo woodhousii'' group of toads (Salientia: Bufonidae). Wasmann Journal of Biology. San Francisco 9: 363–378
*Southwestern Woodhouse's toad – ''Anaxyrus woodhousii australis'' (Shannon & Lowe, 1955)Shannon, F. A., and C. H. Lowe, Jr. 1955. A new subspecies of Bufo woodhousei from the inland Southwest. Herpetologica 11: 185–190.

At one time, Fowler's toad (''Anaxyrus fowleri'') was considered to be a subspecies.

Description

Woodhouse's toad is a robust amphibian and can grow to a maximum snout-vent length of . The head has prominent cranial crests in front of and in between the eyes. The parotoid glands are long and large. The dorsal surface of this toad is grayish-brown or yellowish-brown and it is speckled with small dark spots. There is a narrow pale line running along the spine. The belly is rather paler and is usually unspotted. The male has a single vocal sac on his throat. His call resembles the bleat of a sheep and lasts from one to three seconds.

Distribution and habitat

Woodhouse's toad is found in North America at altitudes of up to . Its range extends from Mexico in the south to Washington in the north. In the United States it is found in Arizona, Arkansas, California, Colorado, Idaho, Kansas, Montana, Nebraska, Nevada, New Mexico, North Dakota, Oklahoma, Oregon, South Dakota, Texas, Utah, Washington, and Wyoming. In the western part of its range it is typically found in lowland riparian corridors, wooded land besides streams and rivers. At higher altitudes it inhabits wet meadows, ponds, reservoirs and lakes. It is also found in urban environments, canals, ponds and irrigated agricultural land.

Behavior

Woodhouse's toad is nocturnal and feeds on insects and other small invertebrates. Near human habitations these toads may congregate underneath outside lights to feed on the insects they attract. Breeding takes place at different times of year in different parts of the range. The males call from in, or close to, standing water and the eggs are laid in gelatinous strings in still-water habitats such as ditches, ponds, pools, cattle tanks and lakes. The tadpoles typically take from five to eight weeks to reach metamorphosis.

Status

Woodhouse's toad has a very wide range and presumed large total population. It is able to live in a number of types of habitat and can tolerate some modification to its habitat. The population seems steady and no particular threats have been identified so the International Union for Conservation of Nature has assessed it as being of "least concern". In central Arizona it seems to be displacing the Arizona toad, (''Anaxyrus microscaphus'').

References

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External links

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Western Soundscape Archive
- Western Woodhouse's Toad recording

{{Taxonbar, from=Q26848327

woodhousii
Amphibians of Mexico
Amphibians of the United States
Taxa named by Charles Frédéric Girard
Amphibians described in 1854