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Wilhelm Friedrich Philipp Pfeffer (9 March 1845 – 31 January 1920) was a German
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
and plant physiologist born in
Grebenstein Grebenstein () is a town in the district of Kassel, in Hesse, Germany. It is located 16 km northwest of Kassel on the German Timber-Frame Road. In 1762 it was the scene of a skirmish between British and French troops during the Seven Years' ...
.


Academic career

He studied
chemistry Chemistry is the scientific study of the properties and behavior of matter. It is a natural science that covers the elements that make up matter to the compounds made of atoms, molecules and ions: their composition, structure, proper ...
and
pharmacy Pharmacy is the science and practice of discovering, producing, preparing, dispensing, reviewing and monitoring medications, aiming to ensure the safe, effective, and affordable use of medication, medicines. It is a miscellaneous science as it ...
at the
University of Göttingen The University of Göttingen, officially the Georg August University of Göttingen, (german: Georg-August-Universität Göttingen, known informally as Georgia Augusta) is a public research university in the city of Göttingen, Germany. Founded ...
, where his instructors included
Friedrich Wöhler Friedrich Wöhler () FRS(For) Hon FRSE (31 July 180023 September 1882) was a German chemist known for his work in inorganic chemistry, being the first to isolate the chemical elements beryllium and yttrium in pure metallic form. He was the fi ...
(1800-1882), William Eduard Weber (1804-1891) and
Wilhelm Rudolph Fittig Wilhelm Rudolph Fittig (6 December 183519 November 1910) was a German chemist. He discovered the pinacol coupling reaction, mesitylene, diacetyl and biphenyl. Fittig studied the action of sodium on ketones and hydrocarbons. He discovered the Fit ...
(1835-1910). Afterwards, he furthered his education at the universities of
Marburg Marburg ( or ) is a university town in the German federal state (''Bundesland'') of Hesse, capital of the Marburg-Biedenkopf district (''Landkreis''). The town area spreads along the valley of the river Lahn and has a population of approximat ...
and
Berlin Berlin ( , ) is the capital and largest city of Germany by both area and population. Its 3.7 million inhabitants make it the European Union's most populous city, according to population within city limits. One of Germany's sixteen constitu ...
. At Berlin, he studied under Alexander Braun (1805-1877) and was an assistant to
Nathanael Pringsheim Nathanael Pringsheim (30 November 1823 – 6 October 1894) was a German botanist. Biography Nathanael Pringsheim was born at Landsberg, Prussian Silesia, and studied at the universities of Breslau, Leipzig, and Berlin successively. He graduat ...
(1823-1894). Later on, he served as an assistant to
Julius von Sachs Julius von Sachs (; 2 October 1832 – 29 May 1897) was a German botanist from Breslau, Prussian Silesia. He is considered the founder of experimental plant physiology and co-founder of modern water culture. Julius von Sachs and Wilhelm Knop a ...
(1832-1897) at
Würzburg Würzburg (; Main-Franconian: ) is a city in the region of Franconia in the north of the German state of Bavaria. Würzburg is the administrative seat of the ''Regierungsbezirk'' Lower Franconia. It spans the banks of the Main River. Würzburg ...
, In 1873 he was appointed professor of
pharmacology Pharmacology is a branch of medicine, biology and pharmaceutical sciences concerned with drug or medication action, where a drug may be defined as any artificial, natural, or endogenous (from within the body) molecule which exerts a biochemica ...
and botany at the
University of Bonn The Rhenish Friedrich Wilhelm University of Bonn (german: Rheinische Friedrich-Wilhelms-Universität Bonn) is a public research university located in Bonn, North Rhine-Westphalia, Germany. It was founded in its present form as the ( en, Rhine ...
, followed by professorships at the Universities of
Basel , french: link=no, Bâlois(e), it, Basilese , neighboring_municipalities= Allschwil (BL), Hégenheim (FR-68), Binningen (BL), Birsfelden (BL), Bottmingen (BL), Huningue (FR-68), Münchenstein (BL), Muttenz (BL), Reinach (BL), Riehen (B ...
(from 1877) and
Tübingen Tübingen (, , Swabian: ''Dibenga'') is a traditional university city in central Baden-Württemberg, Germany. It is situated south of the state capital, Stuttgart, and developed on both sides of the Neckar and Ammer rivers. about one in three ...
(from 1878), where he also served as director of the
Botanischer Garten der Universität Tübingen The Botanischer Garten der Universität Tübingen, also known as the Botanischer Garten Tübingen or the Neuer Botanischer Garten Tübingen, is a botanical garden and arboretum maintained by the University of Tübingen. It is located at Hartmeye ...
. In 1887 he became a professor at the
University of Leipzig Leipzig University (german: Universität Leipzig), in Leipzig in Saxony, Germany, is one of the world's oldest universities and the second-oldest university (by consecutive years of existence) in Germany. The university was founded on 2 December ...
and director of its
botanical garden A botanical garden or botanic gardenThe terms ''botanic'' and ''botanical'' and ''garden'' or ''gardens'' are used more-or-less interchangeably, although the word ''botanic'' is generally reserved for the earlier, more traditional gardens, an ...
. He was elected a member of the
Royal Swedish Academy of Sciences The Royal Swedish Academy of Sciences ( sv, Kungliga Vetenskapsakademien) is one of the royal academies of Sweden. Founded on 2 June 1739, it is an independent, non-governmental scientific organization that takes special responsibility for prom ...
in 1897.


Scientific work

Pfeffer was a pioneer of modern
plant physiology Plant physiology is a subdiscipline of botany concerned with the functioning, or physiology, of plants. Closely related fields include plant morphology (structure of plants), plant ecology (interactions with the environment), phytochemistry (b ...
. His scientific interests included the
thermonastic Nastic movements are non-directional responses to stimuli (e.g. temperature, humidity, light irradiance), and are usually associated with plants. The movement can be due to changes in turgor. Decrease in turgor pressure causes shrinkage while ...
and photonastic movements of flowers, the nyctinastic movements of leaves,
protoplast Protoplast (), is a biological term coined by Hanstein in 1880 to refer to the entire cell, excluding the cell wall. Protoplasts can be generated by stripping the cell wall from plant, bacterial, or fungal cells by mechanical, chemical or e ...
ic physics and
photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in ...
. In 1877, while researching
plant metabolism Plant physiology is a subdiscipline of botany concerned with the functioning, or physiology, of plants. Closely related fields include plant morphology (structure of plants), plant ecology (interactions with the environment), phytochemistry (bio ...
, Pfeffer developed a semi-porous membrane to study the phenomena of
osmosis Osmosis (, ) is the spontaneous net movement or diffusion of solvent molecules through a selectively-permeable membrane from a region of high water potential (region of lower solute concentration) to a region of low water potential (region ...
. The eponymous "Pfeffer cell" is named for the osmometric device he constructed for determining the
osmotic pressure Osmotic pressure is the minimum pressure which needs to be applied to a solution to prevent the inward flow of its pure solvent across a semipermeable membrane. It is also defined as the measure of the tendency of a solution to take in a pure ...
of a solution. During his tenure at Leipzig, Pfeffer published an article on the use of
photography Photography is the art, application, and practice of creating durable images by recording light, either electronically by means of an image sensor, or chemically by means of a light-sensitive material such as photographic film. It is emplo ...
to study plant growth. He wanted to extend the chronophotographic experiments of
Étienne-Jules Marey Étienne-Jules Marey (; 5 March 1830, Beaune, Côte-d'Or – 15 May 1904, Paris) was a French scientist, physiologist and chronophotographer. His work was significant in the development of cardiology, physical instrumentation, aviation, cinema ...
(1830-1904) by producing a short film involving the stages of plant growth. This "movie" would be filmed over a period of weeks by frame-at-a-time exposure taken at regular spaced intervals. Later,
time-lapse photography Time-lapse photography is a technique in which the frequency at which film frames are captured (the frame rate) is much lower than the frequency used to view the sequence. When played at normal speed, time appears to be moving faster and thus ...
would become a commonplace procedure.Who's Who of Victorian Cinema
at www.victorian-cinema.net


Written works

* ''Physiologische Untersuchungen'' - 1873 (Physiological studies) * ''Lehrbuch der Pflanzenphysiologie'' (Textbook of plant physiology). * ''Die periodischen Bewegungen der Blattorgane'' - 1875 (The periodic movements of "leaf organs") * ''Osmotische Untersuchungen – Studien zur Zellmechanik'' - 1877 (Osmotic studies) * ''Beiträge zur Kenntniss der Oxydationsvorgänge in lebenden Zellen'' - 1889 (Contributions to the knowledge of the
oxidative Redox (reduction–oxidation, , ) is a type of chemical reaction in which the oxidation states of substrate change. Oxidation is the loss of electrons or an increase in the oxidation state, while reduction is the gain of electrons or a ...
processes in living cells). * ''Über Aufnahme und Ausgabe ungelöster Körper'' - 1890 * ''Studien zur Energetik der Pflanze'' - 1892 (Studies on the energetics of plants) * ''Druck- und Arbeitsleistung durch wachsende Pflanzen'' - 1893 * ''Untersuchungen über die Entstehung der Schlafbewegungen der Blattorgane'' - 1907 * ''Der Einfluss von mechanischer Hemmung und von Belastung auf die Schlafbewegung'' - 1911 (The influence of mechanical stress on the inhibition of sleep and movement). * ''Beiträge zur Kenntniss der Entstehung der Schlafbewegungen''- 1915 (Contributions to the knowledge on the genesis of sleep movements).


References

* List of publications copied from an equivalent article at the
German Wikipedia The German Wikipedia (german: Deutschsprachige Wikipedia) is the German-language edition of Wikipedia, a free and publicly editable online encyclopedia. Founded on March 16, 2001, it is the second-oldest Wikipedia (after the English Wikipedi ...
. {{DEFAULTSORT:Pfeffer, Wilhelm 1845 births 1920 deaths People from Grebenstein 19th-century German botanists University of Göttingen alumni University of Bonn faculty Members of the Royal Swedish Academy of Sciences Recipients of the Pour le Mérite (civil class) Fellows of the Royal Society Foreign associates of the National Academy of Sciences University of Tübingen faculty University of Basel faculty Leipzig University faculty Plant physiologists People from Mansfeld-Südharz Members of the Royal Society of Sciences in Uppsala