Whatcheeriidae is an extinct

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

tetrapod Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct theraps ...

s which lived in the Mississippian sub-period, a subdivision of the

Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...

period. It contains the genera ''

Pederpes ''Pederpes'' ("Peter's Foot") is an extinct genus of early Carboniferous tetrapod, dating from 348 to 347.6 Ma in the Tournaisian age (lower Mississippian). ''Pederpes'' contains one species, ''P. finneyae'', 1 m long. This most basal Carbonife ...

'', '' Whatcheeria'', and possibly '' Ossinodus''. Fossils of a possible whatcheeriid have been found from the Red Hill locality of

Pennsylvania Pennsylvania (; ( Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, ...

. If these remains are from a whatcheeriid, they extend the range of the family into the

Late Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, wher ...

and suggest that advanced

s may have lived alongside primitive tetrapod ancestors like ''

Hynerpeton ''Hynerpeton'' ( ) is an extinct genus of early four-limbed vertebrate that lived in the rivers and ponds of Pennsylvania during the Late Devonian period, around 365 to 363 million years ago. The only known species of ''Hynerpeton'' is ''H. basse ...

'' and ''

Densignathus ''Densignathus'' is an extinct genus of early stem-tetrapod from the Late Devonian of Pennsylvania in the United States. A lower jaw has been found from the Red Hill fossil site, which is known for a diversity of lobe-finned fishes and other ear ...

''. They also imply that a very long

ghost lineage A ghost lineage is a hypothesized ancestor in a species lineage that has left no fossil evidence yet can be inferred to exist because of gaps in the fossil record or genomic evidence. The process of determining a ghost lineage relies on fossilized ...

of whatcheeriids lived through

Romer's gap Romer's gap is an example of an apparent gap in the tetrapod fossil record used in the study of evolutionary biology. Such gaps represent periods from which excavators have not yet found relevant fossils. Romer's gap is named after paleontologist ...

, a period during the Early Carboniferous conspicuously lacking in tetrapod remains.

Classification

Currently, using modern

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...

taxonomy, Whatcheeriidae is not placed in

Amphibia Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arbore ...

or any other class but simply as its own family within stem-group tetrapods. The analysis below was conducted by Swartz in 2012, showing the relationship of whatcheeriids with other stem-tetrapods.

References

Taxonomicon article on Whatcheeriidae

Zipcode Zoo article on Whatcheeriidae
Carboniferous vertebrates Mississippian animals Prehistoric tetrapod families Stegocephalians Mississippian first appearances Mississippian extinctions {{Carboniferous-animal-stub