Woodward's Wallaroo
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The black wallaroo (''Osphranter bernardus''), also known as Woodward's wallaroo, is a species of macropod restricted to a small, mountainous area in
Arnhem Land Arnhem Land is a historical region of the Northern Territory of Australia. It is located in the north-eastern corner of the territory and is around from the territorial capital, Darwin, Northern Territory, Darwin. In 1623, Dutch East India Compa ...
,
Northern Territory The Northern Territory (abbreviated as NT; known formally as the Northern Territory of Australia and informally as the Territory) is an states and territories of Australia, Australian internal territory in the central and central-northern regi ...
, Australia, between
South Alligator River Alligator Rivers is the name of an area in an Arnhem Land region of the Northern Territory of Australia, containing three rivers, the East, West, and South Alligator Rivers. It is regarded as one of the richest biological regions in Australia, ...
and
Nabarlek The nabarlek (''Petrogale concinna'') is a small species of macropod found in northern Australia. They are a shy and nocturnal animal that resides in rocky hollows and forages in the surrounding area. Their diet is grasses, sedges, and ferns fo ...
. It classified as
near threatened A near-threatened species is a species which has been Conservation status, categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to Endangered species, endangerment in the ne ...
, mostly due to its limited distribution. A large proportion of the range is protected by
Kakadu National Park Kakadu National Park is a protected area in the Northern Territory of Australia, southeast of Darwin. It is a World Heritage Site. Kakadu is also gazetted as a locality, covering the same area as the national park, with 313 people recorded l ...
.


Taxonomy

The description of the species was published by
Walter Rothschild Lionel Walter Rothschild, 2nd Baron Rothschild, Baron de Rothschild, (8 February 1868 – 27 August 1937) was a British banker, politician, zoologist, and soldier, who was a member of the Rothschild family. As a Zionist leader, he was present ...
in 1904. The author initially assigned the species to a new genus as ''Dendrodorcopsis woodwardi'', but revision of new material forwarded to England by the collector John Tunney persuaded the mammalogist
Oldfield Thomas Michael Rogers Oldfield Thomas (21 February 1858 – 16 June 1929) was a British zoologist. Career Thomas worked at the Natural History Museum, London, Natural History Museum on mammals, describing about 2,000 new species and subspecies for ...
that the characteristics of the taxon were assignable to ''
Macropus ''Macropus'', from the Ancient Greek words μάκρος (''mákros''), meaning "long", and πους (''pous''), meaning "foot", is a marsupial genus in the family Macropodidae. It has two extant species of large terrestrial kangaroos. Thirteen kn ...
''. The specific epithet ''woodwardi'' was preoccupied by another subspecies of the genus ('' Macropus robustus woodwardi''), prompting Rothschild to assign the new epithet ''bernardus''. In 2019, a reassessment of macropod taxonomy determined that the species should be moved from the genus ''
Macropus ''Macropus'', from the Ancient Greek words μάκρος (''mákros''), meaning "long", and πους (''pous''), meaning "foot", is a marsupial genus in the family Macropodidae. It has two extant species of large terrestrial kangaroos. Thirteen kn ...
'' to the genus ''
Osphranter ''Osphranter'' is a genus of large marsupials in the family Macropodidae, commonly known as kangaroos and wallaroos (among other species). It contains the largest extant marsupial, the red kangaroo (''O. rufus''). In 2019, a reassessment of mac ...
''. This change was accepted by the
Australian Faunal Directory The Australian Faunal Directory is an online catalogue of taxonomic and biological information on all animal species known to occur within Australia. It is a database produced by the Department of Climate Change, Energy, the Environment and Water ...
in 2020. Other common names of ''O. bernardus'' also include black kangaroo, northern black wallaroo, Bernard's kangaroo, and Bernard's wallaroo.


Description

The black wallaroo is a smaller member of the kangaroos and wallabies, and the smallest of the
wallaroo Wallaroo is a common name for several species of moderately large macropods, intermediate in size between the kangaroos and the wallabies, that are native to Australia and Papua New Guinea. The word "wallaroo" is from the Dharug ''walaru'' wi ...
s. The males' fur colour is predominately very dark brown to black, whereas females are lighter and greyish-brown. The species is identifiable by a body form that is rotund and muscular and ears that are oval in shape and relatively short. The measurements of the head and body combined is up to 730 millimetres, exceeding the tail length of 640 mm or less. The approximate standing height, from the ground to the crown of the head, is 800 mm. The weight range for males is from 19 to 22 kilograms, females are usually around 13 kg. ''Osphranter bernardus'' is by far the smallest of the wallaroos, as well as the most easily distinguished. The species exhibits strong
sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
, the male is uniformly black or dark brown and has yellowish coloration at the forearm, the female is a mid-grey colour and has dark brown to black at the ends of the limbs and tail. The dentition is a diagnostic of ''O. bernardus'', exhibiting a uniquely grooved notch at the second incisor.


Behaviour

The species is extremely shy and will attempt to flee an observer by seeking its refuge at a nearby hill or sandstone
escarpment An escarpment is a steep slope or long cliff that forms as a result of faulting or erosion and separates two relatively level areas having different elevations. Due to the similarity, the term '' scarp'' may mistakenly be incorrectly used inte ...
, and if pursued demonstrates great agility in leaping to ledges or descending to the ground. They also seek respite from the sun beneath escarpments or in the shade of large trees, only venturing away from these to forage for grasses and herbs or access water at a nearby location. Little research has been undertaken on the behaviour of ''O. bernardus'', but it is known to be a shy
nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...
grazer which does not gather in groups. They find refuge from predators and extremes of weather at the rocky escarpments or caves of its habitat and will rapidly seek these whenever they are disturbed. The species remains undercover until emerging at night to forage in the local area. ''Osphranter bernardus'' is not a gregarious species; they are usually seen alone, or as a male and female pair with perhaps one larger young. Individuals may be seen in the company of others of the genus, the tall and slender
antilopine kangaroo The antilopine kangaroo (''Osphranter antilopinus''), also known as the antilopine wallaroo or the antilopine wallaby, is a species of Macropodidae, macropod found in northern Australia: in Cape York Peninsula in Queensland, the Top End of the N ...
(''Osphranter antilopinus'') and the
common wallaroo The common wallaroo (''Osphranter robustus''), also known as the euro, hill wallaroo, or simply wallaroo, is a species of macropod. The word ''euro'' is particularly applied to one subspecies (''O. r. erubescens'').WE Poole and JC Merchant (198 ...
(''Osphranter robustus''), especially when gathering to drink at waterholes.


Distribution and habitat

The distribution range is restricted to area of eastern
Arnhemland Arnhem Land is a historical region of the Northern Territory of Australia. It is located in the north-eastern corner of the territory and is around from the territorial capital, Darwin. In 1623, Dutch East India Company captain Willem Joosten ...
, from the South Aliigator River to Narbalek, and occur within this range at Mt Brockman and
Nourlangie Rock Burrunggui (sometimes spelled Burrunguy, previously called Nourlangie Rock) is located in an outlying sandstone formation of the Arnhem Land Escarpment within the Kakadu National Park in the Northern Territory of Australia. It is the traditional ...
. The favoured habitat has a monsoonal climate, and is most commonly found in these types of woodlands, rainforest over sandstone and on sandy plains dominated by a variety of grasses. The soil of the habitat is very shallow and interspersed with bare rock surfaces, typically sandstone.


In Aboriginal language and culture

In the
Kunwinjku The Kunwinjku (formerly written Gunwinggu) people are an Australian Aboriginal people, one of several groups within the Bininj people, who live around West Arnhem Land to the east of Darwin, Northern Territory. Kunwinjku people generally refer ...
language of West
Arnhem Land Arnhem Land is a historical region of the Northern Territory of Australia. It is located in the north-eastern corner of the territory and is around from the territorial capital, Darwin, Northern Territory, Darwin. In 1623, Dutch East India Compa ...
, the male black wallaroo is known as ' (or ' in the
Kuninjku dialect Kuninjku is a dialect of Bininj Kunwok, an Australian Aboriginal language. The Aboriginal people who speak Kuninjku are the Bininj people, who live primarily in western Arnhem Land. Kuninjku is spoken primarily in the east of the Bininj Kunwok ...
), while the female is called '. It is common in the Kunwinjku language for male and female macropods to have different names. Black wallaroos are said to be the 'pets' of ''Nakidjkidj'' spirits. Kunwinjku sometimes keep them as pets. According to Reverend Peterson Nganjmirra they are 'quiet'.


References


External links


Department of Environment and Heritage Species Profile
{{Taxonbar, from1=Q109262310, from2=Q209585 Macropods Mammals of the Northern Territory Marsupials of Australia Mammals described in 1904 Taxa named by Walter Rothschild