''Woodsia'' is a genus of
ferns in the order
Polypodiales. In the
Pteridophyte Phylogeny Group
The Pteridophyte Phylogeny Group, or PPG, is an informal international group of systematic botanists who collaborate to establish a consensus on the classification of pteridophytes (lycophytes and ferns) that reflects knowledge about plant relation ...
classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder
Aspleniineae.
[ The family can also be treated as the subfamily Woodsioideae of a very broadly defined family ]Aspleniaceae
The Aspleniaceae (spleenworts) are a family of ferns, included in the order Polypodiales. The composition and classification of the family have been subject to considerable changes. In particular, there is a narrow circumscription, Aspleniaceae ...
'' sensu lato''.[ Species of ''Woodsia'' are commonly known as cliff ferns.][
]
Taxonomy
Woodsiaceae formerly included the members of the families Athyriaceae
The Athyriaceae (ladyferns and allies) are a family of terrestrial ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family is placed in the suborder Aspleniineae, and includes two gene ...
and Diplaziopsidaceae
Diplaziopsidaceae is a family of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family is placed in the suborder Aspleniineae, and includes two genera. Alternatively, it may be treat ...
, but analysis has consistently shown that they should be treated as separate families.[ The following cladogram for the suborder Aspleniineae (as eupolypods II), based on Lehtonen (2011),][ and Rothfels & al. (2012),][ shows a likely phylogenetic relationship between the Woodsiaceae and the other families of the Aspleniineae.
]
Species
There are about 40–50 species of the genus ''Woodsia''.[ , '']Plants of the World Online
Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants by ...
'' accepted the following species:[
*'']Woodsia alpina
''Woodsia alpina'', commonly known as alpine woodsia, is a fern found in northern latitudes in North America and Eurasia.[Woodsia andersonii
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...]
'' (Bedd.) Christ
*''Woodsia angolensis
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' Schelpe
*''Woodsia appalachiana
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' T.M.C.Taylor
*''Woodsia asiatica
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' Kiselev & Shmakov
*''Woodsia burgessiana
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' Gerrard ex Hook. & Baker
*''Woodsia calcarea
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' (Fomin) Shmakov
*''Woodsia canescens
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' (Kunze) Mett.
*''Woodsia cinnamomea
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' Christ
*'' Woodsia cochisensis'' Windham
*'' Woodsia cycloloba'' Hand.-Mazz.
*''Woodsia cystopteroides
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' Windham & Mickel
*''Woodsia elongata
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' Hook.
*''Woodsia fragilis
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' (Trevis.) T.Moore
*''Woodsia glabella
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' R.Br. – smooth cliff fern
*''Woodsia gorovoii
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' Krestsch. & Shmakov
*''Woodsia guizhouensis
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' P.S.Wang, Q.Luo & Li Bing Zhang
*''Woodsia hancockii
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' Baker
*''Woodsia heterophylla
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' (Turcz. ex Fomin) Shmakov
*''Woodsia ilvensis
''Woodsia ilvensis'', commonly known as oblong woodsia, is a fern found in North America and northern Eurasia. Also known as rusty woodsia or rusty cliff fern, it is typically found on sunny, exposed cliffs and rocky slopes and on thin, dry, acid ...
'' (L.) R.Br. – oblong woodsia, rusty cliff fern
*''Woodsia indusiosa
''Woodsia indusiosa'' is a species of fern in the family Woodsiaceae. It is endemic to China. Its natural habitat is subtropical or tropical moist lowland forests. It is threatened by habitat loss
Habitat destruction (also termed habitat l ...
'' Christ
*'' Woodsia kangdingensis'' H.S.Kung, Li Bing Zhang & X.S.Guo
*''Woodsia lanosa
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be tre ...
'' Hook.
*'' Woodsia macrochlaena'' Mett. ex Kuhn
*'' Woodsia macrospora'' C.Chr. & Maxon
*'' Woodsia manchuriensis'' Hook.
*'' Woodsia mexicana'' Fée
*'' Woodsia microsora'' Kodama
*'' Woodsia mollis'' (Kaulf.) J.Sm.
*'' Woodsia montevidensis'' Hieron.
*''Woodsia neomexicana
''Physematium neomexicanum'', the New Mexican cliff fern, is a fern species native to the Southwestern United States and northern Mexico.
Distribution
The core of its range is in Coahuila, Nuevo León, Zacatecas, New Mexico, southeastern Utah, A ...
'' Windham – New Mexican cliff fern
*''Woodsia nikkoensis
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be tre ...
'' H.Ogura & Nakaike
*''Woodsia oblonga
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be tre ...
'' Ching & S.H.Wu
*''Woodsia obtusa
''Physematium obtusum'', the bluntlobe cliff fern, is a common rock fern of Appalachia and eastern North America. It prefers a calcareous substrate, but also grows in neutral soils. It may grow on rock faces or in scree.
This fern is often c ...
'' (Spreng.) Torr. – blunt-lobed woodsia, bluntlobe cliff fern
*''Woodsia okamotoi
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' Tagawa
*''Woodsia oregana
''Physematium oreganum'', the Oregon cliff fern, is a perennial fern in the family Woodsiaceae.
This plant is native to a large part of the western and northern United States and Canada
Canada is a country in North America. Its ten p ...
'' D.C.Eaton – Oregon cliff fern
*'' Woodsia phillipsii'' Windham
*'' Woodsia pilosa'' Ching
*''Woodsia pinnatifida
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be tr ...
'' (Fomin) Shmakov
*''Woodsia plummerae
''Physematium plummerae'' is a species of fern known by the common name Plummer's cliff fern. It is native to the southwestern United States and northern Mexico, where it grows in rocky habitat in deserts and other areas. It has leaves up to 25 ce ...
'' Lemmon – Plummer's cliff fern
*'' Woodsia polystichoides'' D.C.Eaton
*''Woodsia pseudoilvensis
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be tre ...
'' Tagawa
*''Woodsia pseudopolystichoides
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be tre ...
'' (Fomin) Kiselev & Shmakov
*'' Woodsia pubescens'' Spreng.
*''Woodsia pulchella
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be tre ...
'' Bertol.
*'' Woodsia rosthorniana'' Diels
*''Woodsia saitoana
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be tre ...
'' Tagawa
*''Woodsia scopulina
''Woodsia scopulina'', common name Rocky Mountain woodsia, is a deciduous perennial fern in the family Woodsiaceae.
This plant is native to the western and northern United States and Canada. ''W. scopulina'' is a small to medium sized fern, 10-3 ...
'' D.C.Eaton – Rocky Mountain woodsia
*''Woodsia shensiensis
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be tre ...
'' Ching
*''Woodsia sinica
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be tre ...
'' Ching
*''Woodsia subcordata
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be tre ...
'' Turcz.
*'' Woodsia subintermedia'' Tzvelev
*''Woodsia taigischensis
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treat ...
'' (Stepanov) Kuznetsov
Some hybrids are also known:[
* ''Woodsia'' × ''abbeae'' Butters
* ''Woodsia'' × ''gracilis'' (G.Lawson) Butters
* ''Woodsia'' × ''kansana'' R.E.Brooks
* ''Woodsia'' × ''maxonii'' R.M.Tryon
* ''Woodsia'' × ''tryonis'' B.Boivin
]
References
{{Taxonbar, from1=Q1314162, from2=Q1633130
Fern genera
Taxonomy articles created by Polbot