The woodcocks are a group of seven or eight very similar living
species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
wading bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s in the
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Scolopax''. The genus name is
Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for a
snipe
A snipe is any of about 26 wading bird species in three genera in the family Scolopacidae. They are characterized by a very long, slender bill, eyes placed high on the head, and cryptic/camouflaging plumage. The ''Gallinago'' snipes have a near ...
or woodcock, and until around 1800 was used to refer to a variety of
wader
245px, A flock of Dunlins and Red knots">Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots
Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflat ...
s.
The English name its first recorded in about 1050.
According to the
Harleian Miscellany
''The Harleian Miscellany'' is a collection of material from the library of the Earl of Oxford and Earl Mortimer collated and edited by Samuel Johnson and William Oldys between 1744 and 1753 on behalf of the publisher Thomas Osborne. Its subtit ...
, a group of woodcocks is called a "fall".
Taxonomy
The
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Scolopax'' was introduced in 1758 by the Swedish naturalist
Carl Linnaeus
Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in the
tenth edition of his ''
Systema Naturae
' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
''. The genus name is
Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for a
snipe
A snipe is any of about 26 wading bird species in three genera in the family Scolopacidae. They are characterized by a very long, slender bill, eyes placed high on the head, and cryptic/camouflaging plumage. The ''Gallinago'' snipes have a near ...
or woodcock.
[ The ]type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is the Eurasian woodcock
The Eurasian woodcock (''Scolopax rusticola'') is a medium-small wading bird found in temperate and subarctic Eurasia. It has cryptic camouflage to suit its woodland habitat, with reddish-brown upperparts and buff-coloured underparts. Its eyes ...
(''Scolopax rusticola'').
Only two woodcocks are widespread, the others being localized island endemics
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
. Most are found in the Northern Hemisphere
The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
but a few range into the Greater Sundas
The Greater Sunda Islands (Indonesian and Malay: ''Kepulauan Sunda Besar'') are four tropical islands situated within Indonesian Archipelago, in the Pacific Ocean. The islands, Borneo, Java, Sulawesi and Sumatra, are internationally recognised fo ...
, Wallacea
Wallacea is a biogeographical designation for a group of mainly Indonesian islands separated by deep-water straits from the Asian and Australian continental shelves. Wallacea includes Sulawesi, the largest island in the group, as well as Lo ...
and New Guinea
New Guinea (; Hiri Motu
Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea).
It is a simplified version of ...
. Their closest relatives are the typical snipe
A snipe is any of about 26 wading bird species in three genera in the family Scolopacidae. They are characterized by a very long, slender bill, eyes placed high on the head, and cryptic/camouflaging plumage. The ''Gallinago'' snipes have a near ...
s of the genus ''Gallinago
''Gallinago'' is a genus of birds in the wader family Scolopacidae, containing 18 species.
Taxonomy
The name ''Gallinago'' was introduced by the French zoologist Mathurin Jacques Brisson in 1760 as a subdivision of the genus ''Scolopax''. Briss ...
''. As with many other sandpiper genera, the lineages that led to ''Gallinago'' and ''Scolopax'' likely diverged around the Eocene
The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
, some 55.8–33.9 million years ago
The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds.
Usage
Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
, although the genus ''Scolopax'' is only known from the late Pliocene
The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58[speciation
Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within ...](_blank)
in island chains, with the extant examples being the Amami woodcock
The Amami woodcock (''Scolopax mira'') is a medium-sized wader. It is slightly larger and longer-legged than Eurasian woodcock, and may be conspecific.
This species is a restricted-range endemic found only in forests on two small islands of the ...
in the Ryukyu Islands
The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Taiwan: the Ōsumi, Tokara, Amami, Okinawa, and Sakishima Islands (further divided into the Miyako and Yaeyama Islands), with Yonaguni ...
and the several species of woodcock in the Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
n islands, the Philippines
The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no),
* bik, Republika kan Filipinas
* ceb, Republika sa Pilipinas
* cbk, República de Filipinas
* hil, Republ ...
, and New Guinea. Subfossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
evidence indicates the presence of another radiation of woodcock species in the Greater Antilles
The Greater Antilles ( es, Grandes Antillas or Antillas Mayores; french: Grandes Antilles; ht, Gwo Zantiy; jam, Grieta hAntiliiz) is a grouping of the larger islands in the Caribbean Sea, including Cuba, Hispaniola, Puerto Rico, Jamaica, and ...
; these Caribbean
The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
woodcocks may have been more closely related to the Old World woodcock species than the New World ones, and were likely wiped out by human incursion into the region.
Species
The genus contains eight species:[
*widespread species
** ]American woodcock
The American woodcock (''Scolopax minor''), sometimes colloquially referred to as the timberdoodle, the bogsucker, the hokumpoke, and the Labrador twister, is a small shorebird species found primarily in the eastern half of North America. Woodcoc ...
, ''Scolopax minor'' (large North America
North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
n range)
** Eurasian woodcock
The Eurasian woodcock (''Scolopax rusticola'') is a medium-small wading bird found in temperate and subarctic Eurasia. It has cryptic camouflage to suit its woodland habitat, with reddish-brown upperparts and buff-coloured underparts. Its eyes ...
, ''Scolopax rusticola'' (large Eurasia
Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
n range)
*localized island endemic species
** Amami woodcock
The Amami woodcock (''Scolopax mira'') is a medium-sized wader. It is slightly larger and longer-legged than Eurasian woodcock, and may be conspecific.
This species is a restricted-range endemic found only in forests on two small islands of the ...
, ''Scolopax mira'' (endemic to the Amami Islands
The The name ''Amami-guntō'' was standardized on February 15, 2010. Prior to that, another name, ''Amami shotō'' (奄美諸島), was also used. is an archipelago in the Satsunan Islands, which is part of the Ryukyu Islands, and is southwest o ...
in Japan
Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
)
** Bukidnon woodcock
The Bukidnon woodcock (''Scolopax bukidnonensis''), or Philippine woodcock is a medium-sized wader. It was only described as new to science as recently as 2001, although the initial specimens had been collected on Luzon in the 1960s, these were ...
, ''Scolopax bukidnonensis'' (endemic to Luzon
Luzon (; ) is the largest and most populous island in the Philippines. Located in the northern portion of the Philippines archipelago, it is the economic and political center of the nation, being home to the country's capital city, Manila, as ...
and Mindanao
Mindanao ( ) ( Jawi: مينداناو) is the second-largest island in the Philippines, after Luzon, and seventh-most populous island in the world. Located in the southern region of the archipelago, the island is part of an island group of ...
in the Philippines
The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no),
* bik, Republika kan Filipinas
* ceb, Republika sa Pilipinas
* cbk, República de Filipinas
* hil, Republ ...
)
** Javan woodcock, ''Scolopax saturata'' (endemic to Sumatra
Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
and Java
Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...
in Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
)
** New Guinea woodcock, ''Scolopax rosenbergii'' (endemic to New Guinea
New Guinea (; Hiri Motu
Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea).
It is a simplified version of ...
)
** Moluccan woodcock, ''Scolopax rochussenii'' (endemic to the Maluku Islands
The Maluku Islands (; Indonesian: ''Kepulauan Maluku'') or the Moluccas () are an archipelago in the east of Indonesia. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone. Geographically they are located eas ...
in Indonesia)
** Sulawesi woodcock
The Sulawesi woodcock (''Scolopax celebensis'') also known as Celebes woodcock, is a medium-sized wader. It is larger and darker than Eurasian woodcock but with small reddish spots.
This species is restricted to wet mountain forests on the Indone ...
, ''Scolopax celebensis'' (endemic to Sulawesi
Sulawesi (), also known as Celebes (), is an island in Indonesia. One of the four Greater Sunda Islands, and the world's eleventh-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Mindanao and the Sulu Ar ...
in Indonesia)
Fossil record
A number of woodcocks are extinct
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
and are known only from fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
or subfossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
bones.
* "Scolopax baranensis" (fossil, Early Pliocene of Hungary; a ''nomen nudum
In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate descr ...
'')
* '' Scolopax carmesinae'' (fossil, Early/Middle Pliocene? of Menorca, Mediterranean)
* '' Scolopax hutchensi'' (fossil, Late Pliocene – Early Pleistocene of Florida, USA)
* ''Scolopax anthonyi
''Scolopax anthonyi'' is a prehistoric species of woodcock in the family Scolopacidae that was once endemic to the Caribbean island of Puerto Rico.
Taxonomy
Its fossil remains were initially identified as belonging to a snipe of the genus ' ...
'' (prehistoric
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of ...
, Holocene of Puerto Rico)[
* '' Scolopax brachycarpa'' (subfossil, Holocene of Hispaniola)
]
Description and ecology
Woodcocks have stocky bodies, cryptic
Cryptic may refer to:
In science:
* Cryptic species complex, a group of species that are very difficult to distinguish from one another
* Crypsis, the ability of animals to blend in to avoid observation
* Cryptic era, earliest period of the Earth
...
brown and blackish plumage, and long slender bills. Their eyes are located on the sides of their heads, which gives them 360° vision. Unlike in most birds, the tip of the bill's upper mandible
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
is flexible.[
As their common name implies, the woodcocks are woodland birds. They feed at night or in the evenings, searching for invertebrates in soft ground with their long bills. This habit and their unobtrusive plumage makes it difficult to see them when they are resting in the day. Most have distinctive displays known as " roding", usually given at dawn or dusk.][
The range of breeding habits of the Eurasian woodcock extends from the west of ]Ireland
Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
eastwards across Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
and Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
preferring mostly boreal forest
Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, ...
regions engulfing northern Japan
Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
, and also from the northern limits of the tree zone in Norway
Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and t ...
. Continuing south to the Pyrenees
The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to C ...
and the northern limits of Spain
, image_flag = Bandera de España.svg
, image_coat = Escudo de España (mazonado).svg
, national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond")
, national_anthem = (English: "Royal March")
, i ...
. Nests have been found in Corsica
Corsica ( , Upper , Southern ; it, Corsica; ; french: Corse ; lij, Còrsega; sc, Còssiga) is an island in the Mediterranean Sea and one of the 18 regions of France. It is the fourth-largest island in the Mediterranean and lies southeast of ...
and there are three isolated Atlantic
The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...
breeding stations in Azores
)
, motto =( en, "Rather die free than subjected in peace")
, anthem= ( en, "Anthem of the Azores")
, image_map=Locator_map_of_Azores_in_EU.svg
, map_alt=Location of the Azores within the European Union
, map_caption=Location of the Azores wi ...
, Madeira
)
, anthem = ( en, "Anthem of the Autonomous Region of Madeira")
, song_type = Regional anthem
, image_map=EU-Portugal_with_Madeira_circled.svg
, map_alt=Location of Madeira
, map_caption=Location of Madeira
, subdivision_type=Sovereign st ...
and the Canary Islands
The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...
. In Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
the sites can be seen as far south as Kashmir
Kashmir () is the northernmost geographical region of the Indian subcontinent. Until the mid-19th century, the term "Kashmir" denoted only the Kashmir Valley between the Great Himalayas and the Pir Panjal Range. Today, the term encompas ...
and the Himalayas
The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
.
Hunting
Some woodcocks have become popular gamebird
Galliformes is an order of heavy-bodied ground-feeding birds that includes turkeys, chickens, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed dispersers and predators, and are often ...
s. The island-endemic species are often quite rare due to overhunting
Overexploitation, also called overharvesting, refers to harvesting a renewable resource to the point of diminishing returns. Continued overexploitation can lead to the destruction of the resource, as it will be unable to replenish. The term app ...
. The pin feathers (coverts of the leading primary feather of the wing) of the Eurasian woodcock are sometimes used as brush
A brush is a common tool with bristles, wire or other filaments. It generally consists of a handle or block to which filaments are affixed in either a parallel or perpendicular orientation, depending on the way the brush is to be gripped durin ...
tips by artist
An artist is a person engaged in an activity related to creating art, practicing the arts, or demonstrating an art. The common usage in both everyday speech and academic discourse refers to a practitioner in the visual arts only. However, th ...
s, by whom they are used for fine painting work.
The cocker spaniel
Cocker Spaniels are dogs belonging to two breeds of the spaniel dog type: the American Cocker Spaniel and the English Cocker Spaniel of which are commonly called simply Cocker Spaniel in their countries of origin. In the early 20th century, Cocker ...
dog breed is named after the bird: the dogs were originally bred to hunt the woodcock.
References
External links
Arthur Cleveland Bent. ''Life Histories of Familiar North American Birds: American Woodcock (Scolopax minor).'' Published in 1927: Smithsonian Institution United States National Museum Bulletin 142 (Part 1): 61–78.
*
{{Taxonbar, from=Q642855
Birds described in 1758
Taxa named by Carl Linnaeus
Wading birds