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William Rowan FRSC (1891–1957), was a Canadian
biologist A biologist is a scientist who conducts research in biology. Biologists are interested in studying life on Earth, whether it is an individual Cell (biology), cell, a multicellular organism, or a Community (ecology), community of Biological inter ...
and
ornithologist Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and th ...
best known as a founder of
photobiology Photobiology is the scientific study of the beneficial and harmful interactions of light (technically, non-ionizing radiation) in living organisms. The field includes the study of photophysics, photochemistry, photosynthesis, photomorphogenesis, vis ...
for establishing the link between the length of daylight and bird migration. Rowan was recognized for his findings with the
Flavelle Medal The Flavelle Medal is an award of the Royal Society of Canada "for an outstanding contribution to biological science during the preceding ten years or for significant additions to a previous outstanding contribution to biological science". It is n ...
in 1946. In 1993, Marianne Gosztonyi Ainley wrote ''Restless Energy: A Biography of William Rowan, 1891–1957'' detailing his life.


Biography

William Rowan was born in
Basel , french: link=no, Bâlois(e), it, Basilese , neighboring_municipalities= Allschwil (BL), Hégenheim (FR-68), Binningen (BL), Birsfelden (BL), Bottmingen (BL), Huningue (FR-68), Münchenstein (BL), Muttenz (BL), Reinach (BL), Riehen (BS ...
,
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
on 29 July 1891. In 1908, he emigrated to Canada and spent three years working as a ranch hand, sketching and photographing wildlife in his spare time, before returning to England and enrolling at
University College London , mottoeng = Let all come who by merit deserve the most reward , established = , type = Public research university , endowment = £143 million (2020) , budget = ...
. In 1914, Rowan's studies were briefly interrupted by the outbreak of the First World War.He briefly joined the London Scottish Rifles, but saw no active service and was discharged on medical grounds in 1915. He graduated in 1917 and after further study gained an MSc in 1919. After a brief spell as a school teacher in England, he was appointed as a lecturer in
Zoology Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
at the
University of Manitoba The University of Manitoba (U of M, UManitoba, or UM) is a Canadian public research university in the province of Manitoba. He moved to
Edmonton Edmonton ( ) is the capital city of the Canadian province of Alberta. Edmonton is situated on the North Saskatchewan River and is the centre of the Edmonton Metropolitan Region, which is surrounded by Alberta's central region. The city ancho ...
in 1920 to found the department of
Zoology Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
at the
University of Alberta The University of Alberta, also known as U of A or UAlberta, is a public research university located in Edmonton, Alberta, Canada. It was founded in 1908 by Alexander Cameron Rutherford,"A Gentleman of Strathcona – Alexander Cameron Rutherfor ...
, heading the department until his retirement in 1956. In addition to being a renowned biologist, Rowan was also known for his love of the arts. He was a self-taught pianist. In addition, he was skilled in the visual arts, especially sculpting, pencil sketching, and photography. His photographic works were featured at the
Royal Photographic Society The Royal Photographic Society of Great Britain, commonly known as the Royal Photographic Society (RPS), is one of the world's oldest photographic societies. It was founded in London, England, in 1853 as the Photographic Society of London with ...
in London. Rowan died in
Edmonton Edmonton ( ) is the capital city of the Canadian province of Alberta. Edmonton is situated on the North Saskatchewan River and is the centre of the Edmonton Metropolitan Region, which is surrounded by Alberta's central region. The city ancho ...
on 30 June 1957.


Bird Migration Experiment

In 1924, Rowan began experimenting with the effects of daylight on
bird migration Bird migration is the regular seasonal movement, often north and south along a flyway, between breeding and wintering grounds. Many species of bird migrate. Migration carries high costs in predation and mortality, including from hunting by ...
. It was generally thought that migration was probably initiated by variations in temperature or barometric pressure. He trapped a number of
dark-eyed junco The dark-eyed junco (''Junco hyemalis'') is a species of junco, a group of small, grayish New World sparrows. This bird is common across much of temperate North America and in summer ranges far into the Arctic. It is a very variable species, much ...
s, which he kept caged, and lengthened the hours of daylight to which they were exposed. This produced spring behaviour in the birds during the middle of winter. This work earned Rowan a doctorate from
University College London , mottoeng = Let all come who by merit deserve the most reward , established = , type = Public research university , endowment = £143 million (2020) , budget = ...
, but a direct link to migration remained to be established: the seasonally disturbed birds appeared ready to fly northwards in spite of the fact that it was still winter; however, it remained to be established where they went after their release. Rowan then carried out the now-legendary experiment that established his reputation. In the autumn of 1931, 500 crows were confined in cages on the south bank of the
North Saskatchewan River The North Saskatchewan River is a glacier-fed river that flows from the Canadian Rockies continental divide east to central Saskatchewan, where it joins with the South Saskatchewan River to make up the Saskatchewan River. Its water flows eventual ...
. The crows were divided into two equally sized groups: an experimental group subjected to artificial light that gradually shortened their nights; and a control group, which experienced natural seasonal reductions in daylight hours. Rowan released the crows in November, after their tails were dyed yellow for purposes of identification. Their release was accompanied by a radio and press campaign offering rewards for every yellow-tailed crow caught or shot. As a result, Rowan was able to account for more than half of the birds. More crows from the control group were accounted for, as they tended either to remain near to the site of their release or to fly south; those from the experimental group behaved as they would in spring, flying north. This identification of the cause of migratory behaviour provided Rowan with international standing as a scientist. Rowan had a long standing correspondence on observations with Elsie Cassels, one of the first female ornithologists in Canada, whose advice and observations he respected.


Accomplishments

* Recipient of
Flavelle Medal The Flavelle Medal is an award of the Royal Society of Canada "for an outstanding contribution to biological science during the preceding ten years or for significant additions to a previous outstanding contribution to biological science". It is n ...
(1946) * Fellow of the
Royal Society of Canada The Royal Society of Canada (RSC; french: Société royale du Canada, SRC), also known as the Academies of Arts, Humanities and Sciences of Canada (French: ''Académies des arts, des lettres et des sciences du Canada''), is the senior national, bil ...
(1934) * Fellow of the
Zoological Society of London The Zoological Society of London (ZSL) is a charity devoted to the worldwide conservation of animals and their habitats. It was founded in 1826. Since 1828, it has maintained the London Zoo, and since 1931 Whipsnade Park. History On 29 ...
* Fellow of the
American Ornithologists Union The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its m ...
(1950) * Member of British Ornithological Union


Publications

*''On Photoperiodism, Reproductive Periodicity, and the Animal Migrations of Birds and Certain Fishes'' (1926) *''Riddle of Migration'' (1931)


References

{{DEFAULTSORT:Rowan, William 1891 births 1957 deaths Alumni of University College London Canadian ornithologists Fellows of the Royal Society of Canada People educated at Bedford School Swiss emigrants to Canada 20th-century Canadian zoologists London Scottish soldiers British Army personnel of World War I