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''Perilla frutescens'', commonly called deulkkae, perilla or Korean perilla, is a species of '' Perilla'' in the mint family
Lamiaceae The Lamiaceae ( ) or Labiatae are a family of flowering plants commonly known as the mint, deadnettle or sage family. Many of the plants are aromatic in all parts and include widely used culinary herbs like basil, mint, rosemary, sage, savory ...
. It is an
annual plant An annual plant is a plant that completes its life cycle, from germination to the production of seeds, within one growing season, and then dies. The length of growing seasons and period in which they take place vary according to geographical ...
native to Southeast Asia and Indian highlands, and is traditionally grown in the Korean peninsula, southern China, Japan and India as a crop. An edible plant, perilla is grown in gardens and attracts butterflies. It is aromatic with a strong mint-like smell. A variety of this plant, ''P. frutescens'' var. ''crispa'' known as " shiso", is widely grown in Japan. In the United States, perilla is a weed pest, toxic to cattle after ingestion.


Names

Along with other plants in the genus '' Perilla'', the plant is commonly called "perilla". It is also referred to as Korean perilla, due to its extensive cultivation in Korea and use in Korean cuisine. In the United States, where the plant has become a weed, the plant is known by many names, such as perilla mint, beefsteak plant, purple perilla, Chinese basil, wild basil, blueweed, Joseph's coat, wild coleus and rattlesnake weed.


Infraspecific taxa

''Perilla frutescens'' has three known varieties. * ''P. frutescens'' (var. ''frutescens'') – called Korean perilla or ''deulkkae''. * ''P. frutescens'' var. ''crispa'' – also called ''shiso'' or ''tía tô''. * ''P. frutescens'' var. ''hirtella'' – also called lemon perilla.


Description

Perilla is an annual plant growing tall, with hairy square stalks. The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are opposite, long and wide, with a broad oval shape, pointy ends, serrated(saw-toothed) margins, and long leafstalks. The leaves are green with occasional touches of purple on the underside. The flowers bloom on
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
s at the end of branches and the main
stalk Stalk or stalking may refer to: Behaviour * Stalk, the stealthy approach (phase) of a predator towards its prey * Stalking, an act of intrusive behaviour or unwanted attention towards a person * Deer stalking, the pursuit of deer for sport Biol ...
in late summer. The
calyx Calyx or calyce (plural "calyces"), from the Latin ''calix'' which itself comes from the Ancient Greek ''κάλυξ'' (''kálux'') meaning "husk" or "pod", may refer to: Biology * Calyx (anatomy), collective name for several cup-like structures ...
, long, consist of upper three sepals and the hairy lower two. The
corolla Corolla may refer to: *Corolla (botany), the petals of a flower, considered as a unit *Toyota Corolla, an automobile model name * Corolla (headgear), an ancient headdress in the form of a circlet or crown * ''Corolla'' (gastropod), a genus of moll ...
is long with its lower lip longer than the upper. Two of the four
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s are long. The fruit is a schizocarp, in diameter, and with reticulate pattern on the outside. Perilla seeds can be soft or hard, being white, grey, brown, and dark brown in colour and globular in shape. 1000 seeds weigh about . Perilla seeds contain about 38-45% lipid. File:Perilla frutescens var. frutescens' flower.jpg, flowers File:Perilla frutescens' raceme.jpg,
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
File:Perilla frutescens var. frutescens' foliage.jpg,
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
File:Ripe deulkkae (Perilla frutescens).jpg, ripe plants (autumn) File:Deulkkae (Perilla frutescens) seeds.jpg, seeds


Cultivation

The plant was introduced into Korea before the Unified Silla era, when it started to be widely cultivated. In its natural state, the yield of perilla leaves and seeds is not high. If the stem is cut about above ground level in summer, a new stalk grows and it produces more fruit. Leaves can be harvested from the stem cut off in the summer, as well as from the new stalk and its branches, throughout summer and autumn. The seeds are harvested in autumn when the fruits are ripe. To collect perilla seeds, the whole plant is harvested and the seeds are beat out of the plant, before being spread for sun drying.


Traditional medicine, phytochemicals, and toxicity

Various perilla varieties are used for traditional medicine in Southeast Asia. Characteristic aroma-active
phytochemical Phytochemicals are chemical compounds produced by plants, generally to help them resist fungi, bacteria and plant virus infections, and also consumption by insects and other animals. The name comes . Some phytochemicals have been used as poisons ...
s in perilla leaves include hydrocarbons,
alcohol Alcohol most commonly refers to: * Alcohol (chemistry), an organic compound in which a hydroxyl group is bound to a carbon atom * Alcohol (drug), an intoxicant found in alcoholic drinks Alcohol may also refer to: Chemicals * Ethanol, one of sev ...
s, aldehydes,
furan Furan is a heterocyclic organic compound, consisting of a five-membered aromatic ring with four carbon atoms and one oxygen atom. Chemical compounds containing such rings are also referred to as furans. Furan is a colorless, flammable, highly ...
s, and
ketone In organic chemistry, a ketone is a functional group with the structure R–C(=O)–R', where R and R' can be a variety of carbon-containing substituents. Ketones contain a carbonyl group –C(=O)– (which contains a carbon-oxygen double bo ...
s, particularly perilla ketone,
egoma ''Perilla frutescens'', commonly called deulkkae, perilla or Korean perilla, is a species of ''Perilla'' in the mint family Lamiaceae. It is an annual plant native to Southeast Asia and Indian highlands, and is traditionally grown in the Kore ...
ketone, and isoegoma ketone. Other compounds include perillaldehyde, limonene, linalool,
beta-caryophyllene Caryophyllene (), more formally (−)-β-caryophyllene, (BCP), is a natural bicyclic sesquiterpene that is a constituent of many essential oils, especially clove oil, the oil from the stems and flowers of '' Syzygium aromaticum'' (cloves), the es ...
,
menthol Menthol is an organic compound, more specifically a monoterpenoid, made synthetically or obtained from the oils of corn mint, peppermint, or other mints. It is a waxy, clear or white crystalline substance, which is solid at room temperature and ...
, and alpha-pinene. The ''crispa'' variety is differentiated by leaf and stem colors, which vary from green to red to purple, indicating the presence of
anthocyanin Anthocyanins (), also called anthocyans, are water-soluble vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart gave the name Anthokyan to a chemical compo ...
s. Although perilla is widely cultivated as an edible plant for humans, it is toxic to cattle and other ruminants, as well as horses. In grazing cattle, plant ketones cause
acute respiratory distress syndrome Acute respiratory distress syndrome (ARDS) is a type of respiratory failure characterized by rapid onset of widespread inflammation in the lungs. Symptoms include shortness of breath (dyspnea), rapid breathing (tachypnea), and bluish skin colo ...
, also called "panting disease".


Adverse effects

Contact dermatitis may occur in people handling the leaves or oil. Consumption of large amounts of seeds has resulted in anaphylaxis.


Nutritional value

Perilla seeds are rich in
dietary fiber Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by the ...
and dietary minerals such as calcium, iron, niacin, protein, and thiamine. Perilla leaves are also rich in vitamins A, C and
riboflavin Riboflavin, also known as vitamin B2, is a vitamin found in food and sold as a dietary supplement. It is essential to the formation of two major coenzymes, flavin mononucleotide and flavin adenine dinucleotide. These coenzymes are involved in ...
.


Use


Culinary


East Asia


= China

= In Manchu cuisine, perilla leaves are used to make ''efen'', ("steamed bun"). The perilla buns are made with glutinous sorghum or
glutinous rice Glutinous rice (''Oryza sativa var. glutinosa''; also called sticky rice, sweet rice or waxy rice) is a type of rice grown mainly in Southeast and East Asia, and the northeastern regions of South Asia, which has opaque grains, very low amylose ...
flour dough filled with
red bean paste Red bean paste () or red bean jam, also called adzuki bean paste or ''anko'' (a Japanese word), is a paste made of red beans (also called "adzuki beans"), used in East Asian cuisine. The paste is prepared by boiling the beans, then mashing or g ...
and wrapped with perilla leaves. The dish is related to Food Exhaustion Day, a traditional Manchu holiday celebrated on every 26th day of the 8th month of the
lunisolar calendar A lunisolar calendar is a calendar in many cultures, combining lunar calendars and solar calendars. The date of Lunisolar calendars therefore indicates both the Moon phase and the time of the solar year, that is the position of the Sun in the Ea ...
.


= Japan

= In Japan, the plant is used far less compared to shiso (''Perilla frutescens'' var. ''crispa''). In the Tōhoku regions of northeastern Japan, it is known as ''jūnen'' ("ten years"), because it was believed to add ten years to a person's lifespan. A local preparation in
Fukushima Prefecture Fukushima Prefecture (; ja, 福島県, Fukushima-ken, ) is a prefecture of Japan located in the Tōhoku region of Honshu. Fukushima Prefecture has a population of 1,810,286 () and has a geographic area of . Fukushima Prefecture borders Miya ...
, called ''shingorō'', consists of half-pounded non-glutinous rice patties, which are skewered, smeared with '' miso'', blended with roasted and ground ''jūnen'' seeds, and roasted over charcoal. Oil pressed from the seeds was historically used to in lamps. The warlord Saitō Dōsan (1494–1556) was said to have been originally a seller of ''egoma'' seed oil.


= Korea

= In
Korean cuisine Korean cuisine has evolved through centuries of social and political change. Originating from ancient agricultural and nomadic traditions in Korea and southern Manchuria, Korean cuisine reflects a complex interaction of the natural envi ...
, ''kkaennip'' or perilla leaves are widely used as a herb and a vegetable. ''Kkaennip'' can be used fresh as a ssam vegetable, fresh or blanched as a namul vegetable, or pickled in soy sauce or soybean paste to make '' jangajji'' (pickle) or kimchi. ''Deulkkae'', the perilla seeds, are either toasted and ground into powder called ''deulkkae-garu'' or toasted and pressed to make perilla oil. Toasted ''deulkkae'' powder is used as a spice and a condiment for '' guk'' (soup), '' namul'' (seasoned vegetable dishes), ''
guksu The Guksu (Korean: 국수전, Hanja: 國手戰) was a Go competition in South Korea. It was held 59 times beginning in 1956, and was discontinued in 2016. Outline The Guksu was a Go competition held by the Hanguk Kiwon, and sponsored by ''The D ...
'' (noodle dishes), kimchi, and '' eomuk'' (fishcake). It is also used as '' gomul'' (coating or topping) for
dessert Dessert is a course (food), course that concludes a meal. The course consists of sweet foods, such as confections, and possibly a beverage such as dessert wine and liqueur. In some parts of the world, such as much of Greece and West Africa, and ...
s: '' Yeot'' and several '' tteok'' (rice cake) varieties can be coated with toasted perilla powder. Perilla oil made from toasted perilla seeds is used as a cooking oil and as a condiment. In Korean-style western food, perilla leaves are sometimes used to substitute basil, and the seed powder and oil is used in salad dressings as well as in
dipping sauce A dip or dipping sauce is a common condiment for many types of food. Dips are used to add flavor or texture to a food, such as pita bread, dumplings, crackers, chopped raw vegetables, fruits, seafood, cubed pieces of meat and cheese, potato chi ...
s. A
Michelin Michelin (; ; full name: ) is a French multinational tyre manufacturing company based in Clermont-Ferrand in the Auvergne-Rhône-Alpes ''région'' of France. It is the second largest tyre manufacturer in the world behind Bridgestone and la ...
-starred restaurant in Seoul serves nutty vanilla ice cream whose secret ingredient is perilla oil. Korean cuisine-Ssam-01.jpg, ''Kkaennip'' (perilla leaves) as a ''ssam'' (wrap) vegetable Kkaennip deulgireum bokkeum.jpg, ''Kkaennip'' stir-fried in perilla oil Kkaennipjeon (perilla).jpg, ''Kkaennip- jeon'' (pan-fried perilla leaves) 깻잎부각.jpg, ''Kkaennip-
bugak ''Bugak'' () is a variety of vegetarian ''twigim'' (deep-fried dish) in Korean cuisine. It is made by deep frying dried vegetables or seaweed coated with ''chapssal-pul'' (; glutinous rice paste) and then drying them again. It is eaten as '' ...
'' (deep-fried perilla leaves) Kkaennip-jangajji 2.jpg, ''Kkaennip- jangajji'' (pickled perilla leaves) Kkaennip-kimchi.jpg, ''Kkaennip- kimchi'' (perilla leaf kimchi) Deulkkae (perilla).jpg, ''Deulkkae'' (perilla seeds) Chueotang chopi (Zanthoxylum piperitum) deulkkae (Perilla frutescens) buchu (Allium tuberosum).jpg, '' Chueo-tang'' (loach soup) served with chopi and ''deulkkae'' (perilla seed) powder Deulkkae-gamja-ongsimi.jpg, '' Gamja-ongsimi'' (potato dough soup) boiled with ''deulkkae'' powder Korean cuisine-Namul-03.jpg, '' Goguma-sun-deulkkae-muchim'' (sweet potato stems seasoned with ''deulkkae'' powder) File:Korean granita.jpg, Perilla '' granita''


South Asia


= India

= In India, perilla seeds are roasted and ground with salt, chilis, and tomatoes to make a savoury side dish or chutney. In Kumaon, the seeds of Bhangira (cultivated Perilla) are eaten raw, the seed oil is used for cooking purposes, and the oil cake is consumed raw or fed to cattle. The roasted seeds are also ground to prepare a spicy chutney. The seeds and leaves of Perilla are also used for flavoring curries in north east India. Manipuri cuisine uses the ground roasted seed in a salad locally known as 'singju'. Known as nei lieh by the Khasis its seeds are used in salads and meat dishes. The Assamese, Bodos and Nagas are also well aware of its uses.


= Nepal

= In Nepal, perilla is called ''silam'' (सिलाम). Perilla seeds are roasted and ground with salt, chilis, and tomatoes to make a savoury dip/side dish or chutney.


Seed oil

Having a distinctive nutty aroma and taste, the oil pressed from the toasted perilla seeds is used as a flavor enhancer, condiment, and a cooking oil in Korean cuisine. The press cake remaining after pressing perilla oil can be used as natural fertilizer or
animal feed Animal feed is food given to domestic animals, especially livestock, in the course of animal husbandry. There are two basic types: fodder and forage. Used alone, the word ''feed'' more often refers to fodder. Animal feed is an important input to ...
.


See also

* '' Shiso'' (''Perilla frutescens'' var. ''crispa'') * ''
Sesame Sesame ( or ; ''Sesamum indicum'') is a flowering plant in the genus ''Sesamum'', also called benne. Numerous wild relatives occur in Africa and a smaller number in India. It is widely naturalized in tropical regions around the world and is cu ...
'' (''Sesamum indicum'')


References

{{Authority control Edible nuts and seeds Herbs Korean cuisine Korean condiments Korean vegetables Lamiaceae