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The white-winged chough (''Corcorax melanorhamphos'') is one of only two surviving members of the Australian mud-nest builders family,
Corcoracidae Corcoracidae is a family of passerine birds known as the Australian mudnesters. The family has sometimes been called Struthideidae in the past; however, despite Struthideidae being an older name than Corcoracidae, the latter name takes precedence ...
, and is the only member of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Corcorax''. It is native to southern and eastern
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
and is an example of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
as it is only distantly related to the European
chough There are two species of passerine birds commonly called chough ( ) that constitute the genus ''Pyrrhocorax'' of the Corvidae (crow) family of birds. These are the red-billed chough (''Pyrrhocorax pyrrhocorax''), and the Alpine chough (or yellow- ...
s that it closely resembles in shape, and for which it was named.


Taxonomy

The white-winged chough was first described by French naturalist
Louis Jean Pierre Vieillot Louis Pierre Vieillot (10 May 1748, Yvetot – 24 August 1830, Sotteville-lès-Rouen) was a French ornithologist. Vieillot is the author of the first scientific descriptions and Linnaean names of a number of birds, including species he collecte ...
in 1817 as ''Coracia melanorhamphos'', other names given include ''Pyrrhocorax leucopterus'' by Dutch zoologist
Coenraad Jacob Temminck Coenraad Jacob Temminck (; 31 March 1778 – 30 January 1858) was a Dutch people, Dutch Aristocracy (class), aristocrat, Zoology, zoologist and museum director. Biography Coenraad Jacob Temminck was born on 31 March 1778 in Amsterdam in the Dut ...
in 1820, and ''Corcorax australis'' by French naturalist René-Primevère Lesson in 1830. before the current name was settled by
Gregory Mathews Gregory Macalister Mathews Order of the British Empire, CBE FRSE FZS FLS (10 September 1876 – 27 March 1949) was an Australian-born amateur ornithologist who spent most of his later life in England. Life He was born in Biamble, New South Wal ...
in 1912. The specific epithet is derived from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
words ''melano-'' 'black' and ''rhamphos'' 'beak'. It is placed in the family known as the mud-nest builders or Corcoracidae, written as Grallinidae in older books before the removal of the genus ''
Grallina ''Grallina'' is a genus of passerine bird native to Australia and New Guinea. It is a member of a group of birds termed monarch flycatchers. Taxonomy Higher taxonomy Long thought to be a member of the mudnest builder family Corcoracidae, the ...
'' to the family Monarchidae. It is one of two remaining species, with the
apostlebird The apostlebird (''Struthidea cinerea''), also known as the grey jumper, lousy jack, happy jack, Donny bird or caw bird is a quick-moving, gray or black bird about 33 cm (13 in) long. It is a native to Australia where it roams woodlands ...
(''Struthidea cinerea''), which differs in appearance but exhibits many behavioural similarities. The mudnest builder family Corcoracidae itself is now placed in a narrower 'Core corvine' group, which contains the
crow A crow is a bird of the genus ''Corvus'', or more broadly a synonym for all of ''Corvus''. Crows are generally black in colour. The word "crow" is used as part of the common name of many species. The related term "raven" is not pinned scientifical ...
s and ravens, shrikes, birds of paradise,
fantail Fantails are small insectivorous songbirds of the genus ''Rhipidura'' in the family Rhipiduridae, native to Australasia, Southeast Asia and the Indian subcontinent. Most of the species are about long, specialist aerial feeders, and named as " ...
s, monarch flycatchers, and
drongo The drongos are a family, Dicruridae, of passerine birds of the Old World tropics. The 30 species in the family are placed in a single genus, ''Dicrurus''. Drongos are mostly black or dark grey, short-legged birds, with an upright stance when ...
s. It is only distantly related to the
red-billed chough The red-billed chough, Cornish chough or simply chough ( ; ''Pyrrhocorax pyrrhocorax''), is a bird in the crow family, one of only two species in the genus '' Pyrrhocorax''. Its eight subspecies breed on mountains and coastal cliffs from the we ...
(''Pyrrhocorax pyrrhocorax'') and
Alpine chough The Alpine chough (), or yellow-billed chough (''Pyrrhocorax graculus'') is a bird in the crow family, one of only two species in the genus '' Pyrrhocorax''. Its two subspecies breed in high mountains from Spain eastwards through southern Europ ...
(''P. graculus''), which are members of the crow family
Corvidae Corvidae is a cosmopolitan family of oscine passerine birds that contains the crows, ravens, rooks, jackdaws, jays, magpies, treepies, choughs, and nutcrackers. In colloquial English, they are known as the crow family or corvids. Currently, 13 ...
. The similarities in appearance of dark
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
and downturned
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
are the result of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
.


Description

White-winged choughs are easily recognised but often mistaken for "crows" (such as the
Australian raven The Australian raven (''Corvus coronoides'') is a passerine bird in the genus ''Corvus'' native to much of southern and northeastern Australia. Measuring in length, it has all-black plumage, beak and mouth, as well as strong grey-black legs and ...
). The white-winged chough is a large, black bird—at about only a little smaller than a raven or a little larger than an
Australian magpie The Australian magpie (''Gymnorhina tibicen'') is a black and white passerine bird native to Australia and southern New Guinea. Although once considered to be three separate species, it is now considered to be one, with nine recognised subs ...
—but has red eyes and a finer, slightly down-curved
beak The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food ...
, similar to a European chough. These red eyes become swollen and brighter in colour when the bird is excited. In flight the large white patches in the wings are immediately obvious, and explain the descriptive part of their common name. Their calls consist primarily of a grating alarm call and a descending piping call. The latter call is diagnostic for the bird in the wild, being significantly different in timbre and melody to that of other birds sharing their
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
.


Behaviour

Flight is a mixture of a slow, deep flapping and short glides: unlike their European namesakes, white-winged choughs are not particularly strong or agile fliers and spend the great majority of their time on the ground, foraging methodically through leaf litter for worms, insects, grain, and snails in a loose group, walking with a distinctive swagger, and calling softly to one another every few seconds. A rich find is the cause of general excitement and all come running in to share in it. The family group walks several kilometres each day through its large territory, foraging as it goes, taking to the air only if disturbed. Choughs are territorial and highly social, living in flocks of from about 4 up to about 20 birds, usually all the offspring of a single pair. Because raising of young is a group effort bands of chough may kidnap fledglings from other family groups so that they will be able to help them to raise their chicks next year.


Feeding

The white-winged chough will
forage Forage is a plant material (mainly plant leaves and stems) eaten by grazing livestock. Historically, the term ''forage'' has meant only plants eaten by the animals directly as pasture, crop residue, or immature cereal crops, but it is also used m ...
in litter and rotten wood for termites and beetles. Although birds have eaten pieces of apple placed out for them on feeding tables, they have not been recorded eating
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
on trees. This species has been observed eating cherry tomatoes, whole, in eastern Australian, Newcastle region. The diet is varied, the white-winged chough eats seeds of various
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
es (
Poaceae Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
), ''
Gahnia grandis ''Gahnia grandis'' is a tussock-forming perennial plant found in southeastern mainland Australia and Tasmania. Originally described by botanist Jacques Labillardière as ''Scleria grandis'' in 1800, it was placed in its current genus by S. T. ...
'', ''
Atriplex ''Atriplex'' () is a plant genus of about 250 species, known by the common names of saltbush and orache (; also spelled orach). It belongs to the subfamily Chenopodioideae of the family Amaranthaceae ''s.l.''. The genus is quite variable and w ...
'', ''
Epacris ''Epacris'' is a genus of about forty species of flowering plants in the family Ericaceae. It was formerly treated in a closely related but separate family Epacridaceae, but the various genera within Epacridaceae including ''Epacris'' have been ...
'', ''
Hibbertia ''Hibbertia'', commonly known as guinea flowers, is a genus of flowering plants in the family Dilleniaceae. They are usually shrubs with simple leaves and usually yellow flowers with five sepals and five petals. There are about 400 species, mo ...
'', ''
Solanum ''Solanum'' is a large and diverse genus of flowering plants, which include three food crops of high economic importance: the potato, the tomato and the eggplant (aubergine, brinjal). It is the largest genus in the nightshade family Solanaceae ...
'', ''
Acacia ''Acacia'', commonly known as the wattles or acacias, is a large genus of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa and Australasia. The genus na ...
'' and ''
Exocarpos ''Exocarpos'' is a genus of flowering shrubs and small trees in the sandalwood family, Santalaceae. They are found throughout Southeast Asia, Australia and the Pacific Islands. They are semi-parasitic, requiring the roots of a host tree, a tr ...
'' species, as well as introduced species such as the cursed thistle (''
Cirsium arvense ''Cirsium arvense'' is a perennial species of flowering plant in the family Asteraceae, native throughout Europe and western Asia, northern Africa and widely introduced elsewhere.Joint Nature Conservation Committee''Cirsium arvense'' The standa ...
''), roadside thistle ('' C. vulgare''), winter honeysuckle ('' Lonicera fragrantissima''), ''
Convolvulus ''Convolvulus'' is a genus of about 200 to 250''Convolvulus''.
Flora of China.
'' and ''
Oxalis ''Oxalis'' ( (American English) or (British English)) is a large genus of flowering plants in the wood-sorrel family Oxalidaceae, comprising over 550 species. The genus occurs throughout most of the world, except for the Polar region, polar are ...
'' species. Berries of the introduced ''
Cotoneaster ''Cotoneaster'' is a genus of flowering plants in the rose family, Rosaceae, native to the Palaearctic region (temperate Asia, Europe, north Africa), with a strong concentration of diversity in the genus in the mountains of southwestern China an ...
'' and ''
Crataegus ''Crataegus'' (), commonly called hawthorn, quickthorn, thornapple, Voss, E. G. 1985. ''Michigan Flora: A guide to the identification and occurrence of the native and naturalized seed-plants of the state. Part II: Dicots (Saururaceae–Cornacea ...
'' are also consumed. It eats a wide variety of arthropods, including
centipede Centipedes (from New Latin , "hundred", and Latin , " foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an ...
s,
millipede Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a resu ...
s and many types of insect—beetles, cockroaches,
termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattode ...
s,
grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are among what is possibly the most ancient living group of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grasshopp ...
s and
crickets Crickets are orthopteran insects which are related to bush crickets, and, more distantly, to grasshoppers. In older literature, such as Imms,Imms AD, rev. Richards OW & Davies RG (1970) ''A General Textbook of Entomology'' 9th Ed. Methuen 8 ...
, flies,
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
and
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s, and
ant Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22 ...
s, bees and
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. Th ...
s. In some areas, it will readily approach tourist sites to find supplementary food.


Breeding

Nesting and breeding is communal, all members of the family helping to raise the young - a process that takes several years, as young birds must learn the art of finding food in the dry Australian bush. Larger families have a better chance of breeding success: so much so that given the opportunity choughs will kidnap the youngsters of neighboring families in order to recruit them to the team. Breeding season is from August to December. The nest is a deep cup-shaped structure made of
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
es held together with mud or sometimes manure in a tree fork up to 10 metres above the ground. Three to five cream-coloured
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
sparsely splotched with dark brown and lavender shades are laid measuring 30 mm x 40 mm. There is one report of white-winged choughs occupying and using a nest which was likely to have been built by the
Australian magpie The Australian magpie (''Gymnorhina tibicen'') is a black and white passerine bird native to Australia and southern New Guinea. Although once considered to be three separate species, it is now considered to be one, with nine recognised subs ...
. However, this was unable to be confirmed as the nest was not witnessed being built. All members of a family take turns to incubate, preen, and feed youngsters, and all cooperate in defending the nest against
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
s. However, the juveniles, who are highly inefficient foragers, have been observed to engage in deception; they bring food back to the
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materia ...
and make to feed nestlings, but instead wait until unobserved, and then eat it themselves. This behaviour disappeared when food sources were artificially supplemented. There are three main threats to young choughs: starvation; predation by nest-robbing birds, particularly
currawong Currawongs are three species of medium-sized passerine birds belonging to the genus ''Strepera'' in the family Artamidae native to Australia. These are the grey currawong (''Strepera versicolor''), pied currawong (''S. graculina''), and black ...
s; and sabotage by neighbouring chough families anxious to protect their food supply by restricting competition. Larger family groups are better able to deal with all three threats. Nesting in communal groups is essential: pairs, or even trios, of birds are almost never successful in raising young. Young take six to seven months to reach independence a significant time-span for birds but they repay the long investment by returning to the nest the next year and helping raise the next generation.


References

David Attenborough, The Life of Birds, Episode 9, 35 mins ff.


External links


White-winged Chough videos, photos & sounds
on the Internet Bird Collection
birdlife.org.au
article
Encyclopedia of Life
article {{Taxonbar, from=Q971196
white-winged chough The white-winged chough (''Corcorax melanorhamphos'') is one of only two surviving members of the Australian mud-nest builders family, Corcoracidae, and is the Monotypic taxon, only member of the genus ''Corcorax''. It is native to southern and e ...
Birds of Queensland Birds of New South Wales Birds of Victoria (Australia) Birds of South Australia Endemic birds of Australia
white-winged chough The white-winged chough (''Corcorax melanorhamphos'') is one of only two surviving members of the Australian mud-nest builders family, Corcoracidae, and is the Monotypic taxon, only member of the genus ''Corcorax''. It is native to southern and e ...
white-winged chough The white-winged chough (''Corcorax melanorhamphos'') is one of only two surviving members of the Australian mud-nest builders family, Corcoracidae, and is the Monotypic taxon, only member of the genus ''Corcorax''. It is native to southern and e ...