White-naped Tit
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The white-naped tit (''Machlolophus nuchalis''), sometimes called white-winged tit, is a
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by th ...
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
in the tit family Paridae. It is endemic to
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
where it is found in dry thorn scrub forest in two disjunct populations, in western India and southern India. Its specific name ''nuchalis'' means ‘of the
nuchal The nape is the back of the neck. In technical anatomical/medical terminology, the nape is also called the nucha (from the Medieval Latin rendering of the Arabic , "spinal marrow"). The corresponding adjective is ''nuchal'', as in the term ''n ...
,
nape The nape is the back of the neck. In technical anatomical/medical terminology, the nape is also called the nucha (from the Medieval Latin rendering of the Arabic , "spinal marrow"). The corresponding adjective is ''nuchal'', as in the term ''nu ...
’. This species is hard to mistake with its contrasting black and white patterns without the grey wing coverts and back of the partly sympatric
cinereous tit The cinereous tit (''Parus cinereus'') is a species of bird in the tit family Paridae. This species is made up of several populations that were earlier treated as subspecies of the great tit (''Parus major''). These birds are grey backed with whi ...
(''Parus cinereus''). This species is very patchily distributed and has been considered to be vulnerable to extinction especially because of the scarcity of suitable habitats particularly nest cavities made by woodpeckers.


Taxonomy

The white-naped tit was formerly one of the many species in the genus ''
Parus ''Parus'' is a genus of Old World birds in the tit family. It was formerly a large genus containing most of the 50 odd species in the family Paridae. The genus was split into several resurrected genera following the publication of a detailed mo ...
'' but was moved to ''
Machlolophus ''Machlolophus'' is a genus of birds in the tit family. The species were formerly placed with many others in the genus ''Parus'' but were moved to ''Machlolophus'' based on a molecular phylogenetic analysis published in 2013 that showed that the ...
'' after a molecular phylogenetic analysis published in 2013 showed that the members of the new genus formed a distinct clade.


Description

The only pied (black-and-white) tit in India, this species has the wing-coverts, crown, sides of head, chin, throat, a ventral band running down the breast and belly to the vent black. The cheeks below the eye, the ear-coverts and a patch on the nape are white. The wing has white on the outer primaries and the base of the secondaries. The last tertiaries are completely white. The two outer tail feathers are white while the next has the outer web white and the remaining black. The white of the flanks can be suffused with yellow.


Distribution and habitat

This species was discovered in the Eastern Ghats near
Nellore Nellore is a city located on the banks of Penna River, in Nellore district of the Indian state of Andhra Pradesh. It serves as the headquarters of the district, as well as Nellore mandal and Nellore revenue division. It is the fourth most p ...
by T C Jerdon who received a specimen from a local hunter. A specimen was later obtained in 1863 from near Bangalore and for a long time the species was not observed anywhere in southern India. A O Hume had suggested in that the two populations might represent different species. The southern population was subsequently noted when
Salim Ali Sálim Moizuddin Abdul Ali (12 November 1896 – 20 June 1987) was an Indian ornithologist and naturalist. Sometimes referred to as the "''Birdman of India''", Salim Ali was the first Indian to conduct systematic bird surveys across Indi ...
collected specimens from the Biligirirangan Hills. The species occurs in the nearby Kaveri valley area where ''Parus cinereus stupae'' is also found. The species has also been reported from the
Chittoor Chittoor is a city and district headquarters in Chittoor district of the Indian state of Andhra Pradesh. It is also the mandal and divisional headquarters of Chittoor mandal and Chittoor revenue division, respectively. The city has a popu ...
district of
Andhra Pradesh Andhra Pradesh (, abbr. AP) is a state in the south-eastern coastal region of India. It is the seventh-largest state by area covering an area of and tenth-most populous state with 49,386,799 inhabitants. It is bordered by Telangana to the ...
. Salim Ali had claimed that the two species were mutually exclusive, however there is no support for this. The distribution in western India is larger and better known, ranging mainly in areas of Kutch and extending into parts of Rajasthan and Haryana. A specimen in the British Museum marked as being from ''Bootan'' (Bhutan) is considered to be in error. Records from Wynaad, Anshi National Park and Dharwad have also been considered doubtful.


Behaviour and ecology

These birds are believed to live at very low densities in small numbers. They are shy and are best detected by their calls which have been described as musical whistling notes ''tee-whi-whi'' or ''see pit-pit-pit-pit''. They forage within well-established territories which pairs will defend from others of their own species. Both males and females make use of roost hollows and individuals have been observed to use the same roost for more than six years. The male of a pair accompanies the female to its roost before proceeds to his own roost site. The birds move their head from side to side in a nearby bush and then fly into the roost cavity, turning around at the entrance so that they can look out and fly if threatened. The birds feed on insects and the berries of ''Salvadora oleoides''. They may also obtain nectar from the flowers of ''Capparis aphylla'' and will sometimes visit rainwater puddles to drink. The breeding season in Rajasthan is during the monsoons, from May to August. The nest is a pad of fibre and hair (sometimes plucked from roadkill) placed inside a cavity typically on a tree. They choose natural hollows as well as cavities made by woodpeckers and coppersmith barbets. Nests have been found in old trees of ''Salvadora persica'' and ''Boswellia serrata''. The clutch size is not known but adults with three young have been observed. The female develops a brood-patch and it is not known if the male incubates. Males have been seen carrying food to the female at nest. Both parents take part in care of the chicks. The chicks are fed mainly with non-hairy caterpillars, the parents may bring food as late as 2000 Hrs in the evening. In Gujarat it has been said to be dependent on old woodpeckers nest holes, especially those made by yellow-crowned woodpeckers, which are used both for nesting and roosting. Suggestions that the species might migrate from western to southern India have not found support and birds (including juveniles) have been seen in southern India during summer. Birds ringed in western India have been recaptured within a radius of 5 to 7 km. Breeding has not been studied in the southern population. This species is said to have undergone a rapid population decline in the recent past. The habitat of dense ''Acacia'' scrub is severely degraded and fragmented in western India especially due to the collection of old branches with potential nest holes for firewood.


References

{{Taxonbar, from=Q27075358 white-naped tit Birds of India Endemic birds of India white-naped tit white-naped tit