Western Sand Darter
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The western sand darter (''Ammocrypta clara'') is a species of freshwater
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...
, a darter from the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Etheostomatinae Etheosomatidae is a species rich subfamily of freshwater ray-finned fish, the members of which are commonly known as the darters. The subfamily is part of the family Percidae which also includes the perches, ruffes and pikeperches. The family i ...
, part of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Percidae The Percidae are a family of ray-finned fish, part of the order Perciformes, which are found in fresh and brackish waters of the Northern Hemisphere. The majority are Nearctic, but there are also Palearctic species. The family contains more than ...
, which also contains the
perch Perch is a common name for fish of the genus ''Perca'', freshwater gamefish belonging to the family Percidae. The perch, of which three species occur in different geographical areas, lend their name to a large order of vertebrates: the Percif ...
es, ruffes and pikeperches. It is native to the central
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
.


Distribution

The western sand darter occurs in river systems from
Lake Michigan Lake Michigan is one of the five Great Lakes of North America. It is the second-largest of the Great Lakes by volume () and the third-largest by surface area (), after Lake Superior and Lake Huron. To the east, its basin is conjoined with that o ...
to
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, including several sections of the
Mississippi Basin The Mississippi River is the second-longest river and chief river of the second-largest drainage system in North America, second only to the Hudson Bay drainage system. From its traditional source of Lake Itasca in northern Minnesota, it fl ...
. Its range extends as far east as the Elk River in
West Virginia West Virginia is a state in the Appalachian, Mid-Atlantic and Southeastern regions of the United States.The Census Bureau and the Association of American Geographers classify West Virginia as part of the Southern United States while the Bur ...
.


Description

This species is up to 7.1 centimeters in length. It is slender and nearly cylindrical in shape. It is pale, translucent silvery white with yellowish coloration along the back. It is distinguished from other sand darters the lack of dark bands or blotches, and by a spine on its operculum.


Biology

This fish lives in medium and large rivers, over sandy and gravel substrates. It requires loose substrate, because it spends much of its time buried in the sand with just its head protruding. This behavior helps it reach cooler temperatures. It feeds on invertebrates, especially the larvae of aquatic insects. It spawns in summer, starting in June in northern regions and May farther south. Females produce an average of 57 eggs at a time, with larger females producing more eggs.


Taxonomy

The Western sand darter was first formally described in 1885 by the
American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish ( Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of Octobe ...
s David Starr Jordan (1851-1931) and Seth Eugene Meek (1859-1914) with the type locality given as the
Des Moines River The Des Moines River () is a tributary of the Mississippi River in the upper Midwestern United States that is approximately long from its farther headwaters.U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe Na ...
at
Ottumwa, Iowa Ottumwa ( ) is a city in and the county seat of Wapello County, Iowa, United States. The population was 25,529 at the time of the 2020 U.S. Census. Located in the state's southeastern section, the city is split into northern and southern halves b ...
. This species forms a
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
with the
naked sand darter The naked sand darter (''Ammocrypta beanii'') is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the Family (biology), family Percidae, which also contains the perches, Gymnocephalus, ruffes and Sande ...
(''A. beanii'') the Florida sand darter (''A. bifascia'').


Conservation

This is considered to be a
vulnerable species A vulnerable species is a species which has been Conservation status, categorized by the International Union for Conservation of Nature as being threatened species, threatened with extinction unless the circumstances that are threatened species, ...
because it has a fragmented distribution and its habitat is degraded in many areas. Increased silt and pollution in river systems reduces the quality of its habitat. The Mississippi River and associated streams and tributaries are heavily channelized and partitioned by
locks Lock(s) may refer to: Common meanings *Lock and key, a mechanical device used to secure items of importance *Lock (water navigation), a device for boats to transit between different levels of water, as in a canal Arts and entertainment * ''Lock ...
and dams, eliminating sites where the fish might live.


References

{{Taxonbar, from=Q2044449


External links


''Ammocrypta clara''.
Nonindigenous Aquatic Species Database. USGS. Ammocrypta Freshwater fish of the United States Fish of the Great Lakes Fish of the Eastern United States Taxa named by David Starr Jordan Taxa named by Seth Eugene Meek Fish described in 1885