Werneria Pygmaea
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''Werneria'', also known as the torrent toads or smalltongue toads, is a small genus of "true toads", family
Bufonidae {{Cat main, Toad This category contains both species commonly called toads, and the true toads from the family Bufonidae {{Cat main, Toad This category contains both species commonly called toads, and the true toads from the family Bufonidae {{Cat m ...
. They are found in western
Central Africa Central Africa is a subregion of the African continent comprising various countries according to different definitions. Angola, Burundi, the Central African Republic, Chad, the Democratic Republic of the Congo, the Republic of the Congo, ...
(Equatorial Guinea, Gabon, and Cameroon), with the greatest
species richness Species richness is the number of different species represented in an ecological community, landscape or region. Species richness is simply a count of species, and it does not take into account the abundances of the species or their relative a ...
in the
Western High Plateau The Western High Plateau, Western Highlands or Bamenda Grassfields is a region of Cameroon characterised by high relief, cool temperatures, heavy rainfall and savanna vegetation. The region lies along the Cameroon line and consists of mounta ...
of
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
. The species generally have restricted or patchy distributions and are considered
threatened Threatened species are any species (including animals, plants and fungi) which are vulnerable to endangerment in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of ''critical depensat ...
.


Etymology

The genus is named after
Franz Werner Franz Josef Maria Werner (15 August 1867 in Vienna – 28 February 1939 in Vienna) was an Austrian zoologist and explorer. Specializing as a herpetologist and entomologist, Werner described numerous species and other taxa of frogs, snakes, inse ...
, Austrian zoologist and herpetologist who was active in Africa.


Description

''Werneria'' are medium-sized toads with a snout-to-vent length between ; females tend to be larger than males. They do not have hearing organs,
vocal sac The vocal sac is the flexible membrane of skin possessed by most male frogs and toads. The purpose of the vocal sac is usually as an amplification of their mating or advertisement call. The presence or development of the vocal sac is one way o ...
s (they are silent), nor
parotoid gland The parotoid gland (alternatively, paratoid gland) is an external skin gland on the back, neck, and shoulder of toads and some frogs and salamanders. It can secrete a number of milky alkaloid substances (depending on the species) known collective ...
s. Skin is smooth. Toe webbing ranges from rudimentary to full.
Tadpole A tadpole is the larval stage in the biological life cycle of an amphibian. Most tadpoles are fully aquatic, though some species of amphibians have tadpoles that are terrestrial. Tadpoles have some fish-like features that may not be found i ...
s have a short, muscular tail and a flat body with a huge sucker mouth.


Species


Habitat

''Werneria'' are associated with torrential forest streams, which also is their breeding habitat. Tadpoles use their sucker mouth to cling to the rocks. Outside the breeding season, adult toads can be found in leaf litter far away from streams. They are known from altitudes between


References

{{Taxonbar, from=Q547143 Amphibians of Sub-Saharan Africa Amphibian genera