Weeverfish
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Weevers (or weeverfish) are nine
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...
species of
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
es of
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Trachinidae, order
Trachiniformes Trachiniformes is an order of percomorph bony fish which is traditionally the suborder Trachinoidei of the Perciformes. However, the classification is also considered as a polyphyly by molecular phylogenies. Trachinidae itself is eventually ...
, part of the
Percomorpha Percomorpha () is a large clade of ray-finned fish that includes the tuna, seahorses, gobies, cichlids, flatfish, wrasse, perches, anglerfish, and pufferfish. Evolution Percomorpha are the most diverse group of teleost fish today. Teleosts, an ...
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
. They are long (up to 37 cm), mainly brown in color, and have venomous spines on their first
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
and
gill A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...
s. During the day, weevers bury themselves in sand, just showing their eyes, and snatch prey as it comes past, which consists of
shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...
and small fish. Weevers are unusual in not having
swim bladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled Organ (anatomy), organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their curren ...
s, as do most
bony fish Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartilag ...
, and as a result, sink as soon as they stop actively swimming. With the exception of '' T. cornutus'' from the southeast Pacific, all species in this family are restricted to the eastern Atlantic (including the Mediterranean). A tenth, extinct species, ''
Callipterus ''Callipterus'' is a genus of aphids in the family Aphididae. ; Names brought to synonymy: ''Callipterus elegans'' Koch, C.l., 1855 and ''Callipterus platani'' Kaltenbach, 1843 are synonyms for ''Tinocallis platani ''Tinocallis platani'' is ...
speciosus'', is known from the
Monte Bolca Monte Bolca is a lagerstätte near Verona, Italy that was one of the first fossil sites with high quality preservation known to Europeans, and is still an important source of fossils from the Eocene. Geology Monte Bolca was uplifted from the T ...
lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...
of the
Lutetian The Lutetian is, in the geologic timescale, a stage or age in the Eocene. It spans the time between . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Midd ...
epoch. Weevers are sometimes used as an ingredient in the recipe for ''
bouillabaisse Bouillabaisse (; oc, bolhabaissa, bullabessa ) is a traditional Provençal fish stew originating in the port city of Marseille. The French and English form ''bouillabaisse'' comes from the Provençal Occitan word ''bolhabaissa'', a compound t ...
''. Weevers are sometimes erroneously called 'weaver fish', although the word is unrelated. In fact, the word 'weever' is believed to derive from the
Old French Old French (, , ; Modern French: ) was the language spoken in most of the northern half of France from approximately the 8th to the 14th centuries. Rather than a unified language, Old French was a linkage of Romance dialects, mutually intelligib ...
word ''wivre'', meaning
serpent Serpent or The Serpent may refer to: * Snake, a carnivorous reptile of the suborder Serpentes Mythology and religion * Sea serpent, a monstrous ocean creature * Serpent (symbolism), the snake in religious rites and mythological contexts * Serp ...
or
dragon A dragon is a reptilian legendary creature that appears in the folklore of many cultures worldwide. Beliefs about dragons vary considerably through regions, but dragons in western cultures since the High Middle Ages have often been depicted as ...
, from the Latin ''
viper The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs tha ...
a''. It is sometimes also known as the viperfish, although it is not related to the
viperfish A viperfish is any species of marine fish in the genus ''Chauliodus''. Viperfishes are mostly found in the mesopelagic zone and are characterized by long, needle-like teeth and hinged lower jaws. A typical viperfish grows to lengths of . Viperf ...
proper (i.e., the
stomiid Stomiidae is a family of deep-sea ray-finned fish, including the barbeled dragonfishes. They are quite small, usually around 15 cm, up to 26 cm. These fish are apex predators and have enormous jaws filled with fang-like teeth. They are ...
s of the genus ''Chauliodus''). In Australia, sand perches of the family
Mugilidae The mullets or grey mullets are a family (Mugilidae) of ray-finned fish found worldwide in coastal temperate and tropical waters, and some species in fresh water. Mullets have served as an important source of food in Mediterranean Europe since R ...
are also known as weevers. In Portugal, the weever is known as ''peixe-aranha'', which translates to 'spider-fish', and in
Catalan Catalan may refer to: Catalonia From, or related to Catalonia: * Catalan language, a Romance language * Catalans, an ethnic group formed by the people from, or with origins in, Northern or southern Catalonia Places * 13178 Catalan, asteroid #1 ...
as ''aranya'', which is identical to the word for 'spider'.


Species

The 9 extant species in two
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
are: * Genus ''
Echiichthys The lesser weever (''Echiichthys vipera'') is a venomous weever of the family Trachinidae, in the order Perciformes, and the class Actinopterygii. It is generally found on the sandy sea beds of the open sea, near the shore. Lesser weevers may st ...
'' **
Lesser weever The lesser weever (''Echiichthys vipera'') is a venomous weever of the family Trachinidae, in the order Perciformes, and the class Actinopterygii. It is generally found on the sandy sea beds of the open sea, near the shore. Lesser weevers may st ...
, ''E. vipera'' ( Cuvier, 1829) * Genus ''
Trachinus ''Trachinus'' is a genus of weevers, order Perciformes that consists of seven extant species. Six of the genus representatives inhabit the waters of Eastern Atlantic Ocean, but only one, ''Trachinus cornutus'', inhabits the South-Eastern Pacific ...
'' ** Spotted weever, ''T. araneus'' Cuvier, 1829 ** Guinean weever, ''T. armatus''
Bleeker Bleeker is a Dutch occupational surname. Bleeker is an old spelling of ''(linnen)bleker'' ("linen bleacher").Sailfin weever, ''T. collignoni'' Roux, 1957 ** '' Trachinus cornutus'' Guichenot, 1848. **
Greater weever The greater weever (''Trachinus draco'', Linnaeus 1758) is a benthic and demersal venomous marine fish of the family Trachinidae. The greater weever is widely distributed along the eastern Atlantic coastline from Norway to Morocco, extending to ...
, ''T. draco''
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1758
**
Striped weever The striped weever, ''Trachinus lineolatus'', is a fish of the family Trachinidae. Widespread in the Eastern Atlantic from Guinea-Bissau to São Tomé Island and Gabon, it is a marine (ocean), marine tropical fish, tropical demersal fish, up to ...
, ''T. lineolatus'' Fischer, 1885 **
Cape Verde weever ''Trachinus pellegrini'', the Cape Verde weever, is a fish of the family Trachinidae. Widespread in the eastern Atlantic along the coasts of Senegal to Nigeria, including the Canary Islands and Cape Verde, and also reported from Mauritania, it i ...
, ''T. pellegrini'' Cadenat, 1937 **
Starry weever The starry weever, or streaked weever, ''Trachinus radiatus'', is a fish of the family Trachinidae widespread in the eastern Atlantic from Gibraltar to the Gulf of Guinea, and probably further south; it is also known from the Mediterranean Sea. ...
, ''T. radiatus'' Cuvier, 1829 * Genus '' Callipterys'' Agassiz 1835 ** Monte Bolca weever, ''C. speciosus'', from the
Lutetian The Lutetian is, in the geologic timescale, a stage or age in the Eocene. It spans the time between . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Midd ...


Timeline

ImageSize = width:1000px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:10px Period = from:-65.5 till:10 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:5 start:-65.5 ScaleMinor = unit:year increment:1 start:-65.5 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:cenozoic value:rgb(0.54,0.54,0.258) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:neogene value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: -65.5 till: -55.8 color:paleocene text:
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
from: -55.8 till: -33.9 color:eocene text:
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
from: -33.9 till: -23.03 color:oligocene text:
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
from: -23.03 till: -5.332 color:miocene text:
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text:
Pleist. The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
from: -0.0117 till: 0 color:holocene text: H. bar:eratop from: -65.5 till: -23.03 color:paleogene text:
Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...
from: -23.03 till: -2.588 color:neogene text:
Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...
from: -2.588 till: 0 color:quaternary text: Q. PlotData= align:left fontsize:M mark:(line,white) width:5 anchor:till align:left color:eocene bar:NAM1 from: -33.9 till: 0 text:
Trachinus ''Trachinus'' is a genus of weevers, order Perciformes that consists of seven extant species. Six of the genus representatives inhabit the waters of Eastern Atlantic Ocean, but only one, ''Trachinus cornutus'', inhabits the South-Eastern Pacific ...
PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 bar:period from: -65.5 till: -55.8 color:paleocene text:
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
from: -55.8 till: -33.9 color:eocene text:
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
from: -33.9 till: -23.03 color:oligocene text:
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
from: -23.03 till: -5.332 color:miocene text:
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text:
Pleist. The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
from: -0.0117 till: 0 color:holocene text: H. bar:era from: -65.5 till: -23.03 color:paleogene text:
Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...
from: -23.03 till: -2.588 color:neogene text:
Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...
from: -2.588 till: 0 color:quaternary text: Q.


Interaction with humans


Stings: causes, frequency and prevention

Most human stings are inflicted by the lesser weever, which habitually remains buried in sandy areas of shallow water and is thus more likely to come into contact with bathers than other species (such as the greater weever, which prefers deeper water); stings from other species are generally limited to anglers and commercial fishermen. Even very shallow water (sometimes little more than damp sand) may harbour lesser weevers. The vast majority of injuries occur to the foot and are the result of stepping on buried fish; other common sites of injury are the hands and buttocks. Stings are most common in the hours before and after low tide (especially at springs), so one possible precaution is to avoid bathing or paddling at these times. Weever stings have been known to penetrate
wet suit A wetsuit is a garment worn to provide thermal protection while wet. It is usually made of foamed neoprene, and is worn by surfers, divers, windsurfers, canoeists, and others engaged in water sports and other activities in or on water. Its pu ...
boots even through a
rubber Rubber, also called India rubber, latex, Amazonian rubber, ''caucho'', or ''caoutchouc'', as initially produced, consists of polymers of the organic compound isoprene, with minor impurities of other organic compounds. Thailand, Malaysia, and ...
sole (if thin), and bathers and surfers should wear sandals, "
jelly shoes Jelly shoes or jellies are a type of shoe made of PVC plastic. Jelly shoes come in a large variety of brands and colours and the material is sometimes infused with glitter. Its name comes from the French company called Jelly Shoes, founded by Ton ...
", or wetsuit boots with relatively hard soles, and avoid sitting or "rolling" in the shallows. Stings also increase in frequency during the summer (to a maximum in August), but this is probably the result of the greater number of bathers. The lesser weever can be found from the southern
North Sea The North Sea lies between Great Britain, Norway, Denmark, Germany, the Netherlands and Belgium. An epeiric sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian S ...
to the Mediterranean, and is common around the south coast of the United Kingdom and Ireland, the Atlantic coast of France, Portugal and Spain, and the northern coast of the Mediterranean. The high number of bathers found on popular tourist beaches in these areas means stings are common, although individual chances of being stung are low. The ''
South Wales Evening Post The ''South Wales Evening Post'' is a tabloid daily newspaper distributed in the South West region of Wales. The paper has three daily editions – Swansea, Neath and Port Talbot and Carmarthenshire – and is published by Media Wales, part ...
'' stated (on 8 August 2000) that around 40 weever stings are recorded in the
Swansea Swansea (; cy, Abertawe ) is a coastal city and the second-largest city of Wales. It forms a principal area, officially known as the City and County of Swansea ( cy, links=no, Dinas a Sir Abertawe). The city is the twenty-fifth largest in ...
and
Gower Gower ( cy, Gŵyr) or the Gower Peninsula () in southwest Wales, projects towards the Bristol Channel. It is the most westerly part of the historic county of Glamorgan. In 1956, the majority of Gower became the first area in the United Kingdom ...
area every year, but many victims do not seek medical assistance and go uncounted.


Symptoms

At first, many victims believe they have simply scratched themselves on a sharp stone or shell, although this barely hurts; significant pain begins about 2–3 minutes after being stung. Weever stings cause severe pain; common descriptions from victims are "extremely painful" and "much worse than a
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. Th ...
(or bee) sting". Common and minor symptoms include severe pain, itching, swelling, heat, redness, numbness, tingling, nausea, vomiting, joint aches, headaches, abdominal cramps, lightheadedness, increased urination, and tremors. Rare and severe symptoms include abnormal heart rhythms, weakness, shortness of breath, seizures, decreased blood pressure,
gangrene Gangrene is a type of tissue death caused by a lack of blood supply. Symptoms may include a change in skin color to red or black, numbness, swelling, pain, skin breakdown, and coolness. The feet and hands are most commonly affected. If the ga ...
, tissue degeneration,Branko Šuljić, ''Sportski ribolov'', 2001 and unconsciousness.


Treatment

Although extremely unpleasant, weever stings are not generally dangerous and the pain will ease considerably within a few hours even if untreated. Complete recovery may take a week or more; in a few cases, victims have reported swelling and/or stiffness persisting for months after envenomation.
First aid First aid is the first and immediate assistance given to any person with either a minor or serious illness or injury, with care provided to preserve life, prevent the condition from worsening, or to promote recovery. It includes initial in ...
treatment consists of immersing the affected area in hot water (as hot as the victim can tolerate without being scalded), which will accelerate denaturation of the
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
-based
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
. The use of hot water will reduce the pain felt by the victim after a few minutes. Usual experience is that the pain then fades within 10 to 20 minutes, as the water cools.
Folklore Folklore is shared by a particular group of people; it encompasses the traditions common to that culture, subculture or group. This includes oral traditions such as tales, legends, proverbs and jokes. They include material culture, ranging ...
often suggests the addition of substances to the hot water, including
urine Urine is a liquid by-product of metabolism in humans and in many other animals. Urine flows from the kidneys through the ureters to the urinary bladder. Urination results in urine being excretion, excreted from the body through the urethra. Cel ...
,
vinegar Vinegar is an aqueous solution of acetic acid and trace compounds that may include flavorings. Vinegar typically contains 5–8% acetic acid by volume. Usually, the acetic acid is produced by a double fermentation, converting simple sugars to et ...
, and
Epsom salts Magnesium sulfate or magnesium sulphate (in English-speaking countries other than the US) is a chemical compound, a salt with the formula , consisting of magnesium cations (20.19% by mass) and sulfate anions . It is a white crystalline solid, s ...
, but this is of limited (if any) value. Heat should be applied for at least 15 minutes, but the longer the delay (before heat is applied), the longer the treatment should be continued. Once the pain has eased, the injury should be checked for the remains of broken spines, and any found need to be removed. Over-the-counter
analgesic An analgesic drug, also called simply an analgesic (American English), analgaesic (British English), pain reliever, or painkiller, is any member of the group of drugs used to achieve relief from pain (that is, analgesia or pain management). It ...
s, such as
aspirin Aspirin, also known as acetylsalicylic acid (ASA), is a nonsteroidal anti-inflammatory drug (NSAID) used to reduce pain, fever, and/or inflammation, and as an antithrombotic. Specific inflammatory conditions which aspirin is used to treat inc ...
or
ibuprofen Ibuprofen is a nonsteroidal anti-inflammatory drug (NSAID) that is used for treating pain, fever, and inflammation. This includes painful menstrual periods, migraines, and rheumatoid arthritis. It may also be used to close a patent ductus arte ...
, may be of assistance in management of pain and can also reduce
edema Edema, also spelled oedema, and also known as fluid retention, dropsy, hydropsy and swelling, is the build-up of fluid in the body's Tissue (biology), tissue. Most commonly, the legs or arms are affected. Symptoms may include skin which feels t ...
. Medical advice should be sought if any of the symptoms listed above as rare or severe are observed, if swelling spreads beyond the immediate area of injury (e.g. from hand to arm), if symptoms persist, or if any other factor causes concern. Medical treatment consists of symptom management, analgesia (often with
opiate An opiate, in classical pharmacology, is a substance derived from opium. In more modern usage, the term ''opioid'' is used to designate all substances, both natural and synthetic, that bind to opioid receptors in the brain (including antagonis ...
s) and the same heat treatment as for first aid - more systemic treatment using
histamine antagonist Antihistamines are drugs which treat allergic rhinitis, common cold, influenza, and other allergies. Typically, people take antihistamines as an inexpensive, generic (not patented) drug that can be bought without a prescription and provides re ...
s may assist in reducing local inflammation.


Fatalities

The only recorded death in the UK occurred in 1933, when a fisherman off
Dungeness Dungeness () is a headland on the coast of Kent, England, formed largely of a shingle beach in the form of a cuspate foreland. It shelters a large area of low-lying land, Romney Marsh. Dungeness spans Dungeness Nuclear Power Station, the hamlet ...
suffered multiple stings. The victim may have died of other medical causes exacerbated by the stings.


References


External links


Personal accounts of being stung by Weever fish
{{Taxonbar, from=Q649808 * Trachiniformes Venomous fish Taxa named by Constantine Samuel Rafinesque