The waxwings are three species of
passerine
A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s classified in the
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Bombycilla''. They are pinkish-brown and pale grey with distinctive smooth plumage in which many body feathers are not individually visible, a black and white eyestripe, a crest, a square-cut tail and pointed wings. Some of the wing feathers have red tips, the resemblance of which to
sealing wax
Sealing wax is a wax material of a seal which, after melting, hardens quickly (to paper, parchment, ribbons and wire, and other material) forming a bond that is difficult to separate without noticeable tampering. Wax is used to verify something su ...
gives these birds their common name. According to most authorities, this is the only genus placed in the family Bombycillidae, although sometimes the family is extended to include related taxa that are more usually included in separate families: silky flycatchers (
Ptiliogonatidae
The silky-flycatchers are a small family, Ptiliogonatidae, of passerine birds. The family contains only four species in three genera. They were formerly lumped with waxwings and hypocolius in the family Bombycillidae, and they are listed in that ...
(e.g. ''
Phainoptila'')), ''Hypocolius'' (
Hypocoliidae
The grey hypocolius or simply hypocolius (''Hypocolius ampelinus'') is a small passerine bird species. It is the sole member of the genus ''Hypocolius'' and it is placed in a family of its own, the Hypocoliidae. This slender and long tailed bird i ...
), ''Hylocitrea'' (
Hylocitreidae
The hylocitrea (''Hylocitrea bonensis''), also known as the yellow-flanked whistler or olive-flanked whistler, is a species of bird that is endemic to montane forests on the Indonesian island of Sulawesi.Boles, W. E. (2007). Yellow-flanked Whist ...
), palmchats (
Dulidae
The palmchat (''Dulus dominicus'') is a small, long-tailed passerine bird, the only species in the genus ''Dulus'' and the family Dulidae endemic to the Caribbean island of Hispaniola (split between the Dominican Republic and Haiti). It is r ...
) and the Hawai'ian 'honeyeaters' (
Mohoidae
Mohoidae, also known as the Hawaiian honeyeaters, is a family of Hawaiian species of recently extinct, nectarivorous songbirds in the genera '' Moho'' (ōō) and '' Chaetoptila'' (kioea). These now extinct birds form their own family, represent ...
). There are three species: the
Bohemian waxwing
The Bohemian waxwing (''Bombycilla garrulus'') is a starling-sized passerine bird that breeds in the northern forests of the Palearctic and North America. It has mainly buff-grey plumage, black face markings and a pointed crest. Its wings are ...
(''B. garrulus''), the
Japanese waxwing
The Japanese waxwing (''Bombycilla japonica'') is a fairly small passerine bird of the waxwing family found in the East Palaearctic. It feeds mainly on fruit and berries but also eats insects during the summer. The nest is a cup of twigs lined wi ...
(''B. japonica'') and the
cedar waxwing
The cedar waxwing (''Bombycilla cedrorum'') is a member of the family Bombycillidae or waxwing family of passerine birds. It is a medium-sized, mostly brown, gray, and yellow. This bird is named for its wax-like wing tips. It is a native of Nort ...
(''B. cedrorum'').
Waxwings are not long-distance migrants, but move nomadically outside the breeding season. Waxwings mostly feed on fruit, but at times of year when fruits are unavailable they feed on sap, buds, flowers and insects. They catch insects by gleaning through foliage or in mid-air. They often nest near water, the female building a loose nest at the fork of a branch, well away from the trunk of the tree. She also incubates the eggs, the male bringing her food to the nest, and both sexes help rear the young. Waxwings appear in art and have been mentioned in literature.
Etymology
''Bombycilla'', the genus name, is
Vieillot
Louis Pierre Vieillot (10 May 1748, Yvetot – 24 August 1830, Sotteville-lès-Rouen) was a French ornithologist.
Vieillot is the author of the first scientific descriptions and Linnaean names of a number of birds, including species he collect ...
's attempt at
Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for "silktail", translating the German name ''Seidenschwänze''. Vieillot analyzed ''motacilla'', Latin for
wagtail
Wagtails are a group of passerine birds that form the genus ''Motacilla'' in the family Motacillidae. The forest wagtail belongs to the monotypic genus ''Dendronanthus'' which is closely related to ''Motacilla'' and sometimes included therein. ...
, as ''mota'' for "move" and ''cilla'', which he thought meant "tail"; however, ''Motacilla'' actually combines ''motacis'', a mover, with the
diminutive
A diminutive is a root word that has been modified to convey a slighter degree of its root meaning, either to convey the smallness of the object or quality named, or to convey a sense of intimacy or endearment. A (abbreviated ) is a word-formati ...
suffix ''-illa''. He then combined this "''cilla"'' with the Latin ''bombyx'', meaning silk.
Description
Waxwings are characterised by soft silky plumage. They have unique red tips to some of the wing feathers where the shafts extend beyond the barbs; in the
Bohemian
Bohemian or Bohemians may refer to:
*Anything of or relating to Bohemia
Beer
* National Bohemian, a brand brewed by Pabst
* Bohemian, a brand of beer brewed by Molson Coors
Culture and arts
* Bohemianism, an unconventional lifestyle, origin ...
and
cedar waxwing
The cedar waxwing (''Bombycilla cedrorum'') is a member of the family Bombycillidae or waxwing family of passerine birds. It is a medium-sized, mostly brown, gray, and yellow. This bird is named for its wax-like wing tips. It is a native of Nort ...
s, these tips look like sealing wax, and give the group its common name.
The legs are short and strong, and the wings are pointed. The male and female have the same plumage. All three species have mainly brown plumage, a black line through the eye and black under the chin, a square-ended tail with a red or yellow tip, and a pointed crest. The bill, eyes, and feet are dark. The adults moult between August and November, but may suspend their moult and continue after migration. Calls are high-pitched, buzzing or trilling monosyllables.
Behavior
Diet
These are arboreal birds that breed in northern forests.
Their main food is fruit, which they eat from early summer (
strawberries
The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid species of the genus '' Fragaria'', collectively known as the strawberries, which are cultivated worldwide for their fruit. The fruit is widely ap ...
,
mulberries
''Morus'', a genus of flowering plants in the family Moraceae, consists of diverse species of deciduous trees commonly known as mulberries, growing wild and under cultivation in many temperate world regions. Generally, the genus has 64 identif ...
, and
serviceberries) through late summer and fall (
raspberries
The raspberry is the edible fruit of a multitude of plant species in the genus ''Rubus'' of the rose family, most of which are in the subgenus '' Idaeobatus''. The name also applies to these plants themselves. Raspberries are perennial with ...
,
blackberries
The blackberry is an edible fruit produced by many species in the genus ''Rubus'' in the family (biology), family Rosaceae, hybrids among these species within the subgenus ''Rubus'', and hybrids between the subgenera ''Rubus'' and ''Idaeobatus' ...
,
cherries, and
honeysuckle
Honeysuckles are arching shrubs or twining vines in the genus ''Lonicera'' () of the family Caprifoliaceae, native to northern latitudes in North America and Eurasia. Approximately 180 species of honeysuckle have been identified in both conti ...
berries) into late fall and winter (
juniper
Junipers are coniferous trees and shrubs in the genus ''Juniperus'' () of the cypress family Cupressaceae. Depending on the taxonomy, between 50 and 67 species of junipers are widely distributed throughout the Northern Hemisphere, from the Arcti ...
berries,
grape
A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus ''Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters.
The cultivation of grapes began perhaps 8,000 years ago, ...
s,
crabapple
''Malus'' ( or ) is a genus of about 30–55 species of small deciduous trees or shrubs in the family Rosaceae, including the domesticated orchard apple, crab apples, wild apples, and rainberries.
The genus is native to the temperate zone o ...
s,
mountain-ash fruits,
rose
A rose is either a woody perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred species and tens of thousands of cultivars. They form a group of plants that can be ...
hips,
cotoneaster
''Cotoneaster'' is a genus of flowering plants in the rose family, Rosaceae, native to the Palaearctic region (temperate Asia, Europe, north Africa), with a strong concentration of diversity in the genus in the mountains of southwestern China an ...
fruits,
dogwood
''Cornus'' is a genus of about 30–60 species of woody plants in the family Cornaceae, commonly known as dogwoods, which can generally be distinguished by their blossoms, berries, and distinctive bark. Most are deciduous trees or shrub ...
berries, and
mistletoe
Mistletoe is the common name for obligate hemiparasitic plants in the order Santalales. They are attached to their host tree or shrub by a structure called the haustorium, through which they extract water and nutrients from the host plant.
...
berries).
They pluck fruit from a perch or occasionally while hovering. In spring they replace fruit with sap, buds, and flowers. In the warmer part of the year they catch many insects by
gleaning
Gleaning is the act of collecting leftover crops from farmers' fields after they have been commercially harvested or on fields where it is not economically profitable to harvest. It is a practice described in the Hebrew Bible that became a legall ...
or
in midair, and often nest near water where flying insects are abundant.
Reproduction
Waxwings also choose nest sites in places with rich supplies of fruit and breed late in the year to take advantage of summer ripening. However, they may start courting as early as the winter. Pairing includes a ritual in which mates pass a fruit or small inedible object back and forth several times until one eats it (if it is a fruit). After this they may copulate. So that many birds can nest in places with good food supplies, a pair does not defend a territory—perhaps the reason waxwings have no true
song
A song is a musical composition intended to be performed by the human voice. This is often done at distinct and fixed pitches (melodies) using patterns of sound and silence. Songs contain various forms, such as those including the repetitio ...
—but a bird may attack intruders, perhaps to guard its mate. Both birds gather nest materials, but the female does most of the construction, usually on a horizontal limb or in a crotch well away from the tree trunk, at any height. She makes a loose, bulky nest of twigs, grass, and
lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.[pine
A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...]
needles and may camouflage with dangling pieces of grass, flowers, lichen, and moss. The female incubates, fed by the male on the nest, but once the eggs hatch, both birds feed the young.
File:Cedar waxwing Courtship.jpg, Cedar waxwing pair passing a berry back and forth during courtship
File:Cedar Waxwing nest and eggs (14723635678).jpg, Cedar waxwing nest and eggs
Migration
They are not true long-distance
migrants, but wander erratically outside the breeding season and move south from their summer range in winter. In poor berry years huge numbers can irrupt well beyond their normal range, often in flocks that on occasion number in the thousands.
Taxonomy
Some authorities (including the Sibley-Monroe checklist) place some other genera in the family Bombycillidae along with the waxwings. Birds that are sometimes classified in this way include the
silky-flycatcher
The silky-flycatchers are a small family, Ptiliogonatidae, of passerine birds. The family contains only four species in three genera. They were formerly lumped with waxwings and hypocolius in the family Bombycillidae, and they are listed in tha ...
s, the
hypocolius, and the
palm chat
The palmchat (''Dulus dominicus'') is a small, long-tailed passerine bird, the Monotypic taxon, only species in the genus ''Dulus'' and the family Dulidae endemic to the Caribbean island of Hispaniola (split between the Dominican Republic and Hai ...
. Recent molecular analyses have corroborated their affinity and identified them as a
clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
, identifying the
yellow-flanked whistler as another member.
Species
In popular culture
Waxwings are mentioned in the first lines of the poem "Pale Fire" by "
John Shade
''Pale Fire'' is a 1962 novel by Vladimir Nabokov. The novel is presented as a 999-line poem titled "Pale Fire", written by the fictional poet John Shade, with a foreword, lengthy commentary and index written by Shade's neighbor and academic col ...
", a fictional poet created by
Vladimir Nabokov
Vladimir Vladimirovich Nabokov (russian: link=no, Владимир Владимирович Набоков ; 2 July 1977), also known by the pen name Vladimir Sirin (), was a Russian-American novelist, poet, translator, and entomologist. Bo ...
for his novel ''
Pale Fire
''Pale Fire'' is a 1962 novel by Vladimir Nabokov. The novel is presented as a 999-line poem titled "Pale Fire", written by the fictional poet John Shade, with a foreword, lengthy commentary and index written by Shade's neighbor and academic co ...
''.
The waxwing identified in the Commentary to ''Pale Fire'' is a fictitious species. The novel's narrator claims that John Shade's father had a waxwing named for him, ''Bombycilla Shadei'', and in noting the name corrects the taxonomical error: '(this should be ''shadei'', of course)'.
The line "I was the shadow of the waxwing slain" was also used in the song "The Obituaries" by the band
The Menzingers
The Menzingers are an American punk rock band from Scranton, Pennsylvania, formed in 2006. The band consists of Greg Barnett (vocals, guitar), Tom May (vocals, guitar), Eric Keen (bass) and Joe Godino (drums). To date, the band has released se ...
.
Waxwings are also mentioned in the song "Autumn" by
Joanna Newsom.
References
External links
*
*
Waxwing videoson the Internet Bird Collection
{{Authority control
Extant Miocene first appearances
Taxa named by Louis Jean Pierre Vieillot