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The waved albatross (''Phoebastria irrorata''), also known as Galapagos albatross,Remsen Jr., J.V. (2008) is the only member of the family
Diomedeidae Albatrosses, of the biological family Diomedeidae, are large seabirds related to the procellariids, storm petrels, and diving petrels in the order Procellariiformes (the tubenoses). They range widely in the Southern Ocean and the North Pacif ...
located in the tropics. When they forage, they follow a straight path to a single site off the coast of
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
, about to the east. During the non-breeding season, these birds reside primarily on the
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ' ...
ian and
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
vian coasts.


Taxonomy

Waved albatrosses are a species of albatross belonging to family
Diomedeidae Albatrosses, of the biological family Diomedeidae, are large seabirds related to the procellariids, storm petrels, and diving petrels in the order Procellariiformes (the tubenoses). They range widely in the Southern Ocean and the North Pacif ...
of the order
Procellariiformes Procellariiformes is an order (biology), order of seabirds that comprises four family (biology), families: the albatrosses, the Procellariidae, petrels and shearwaters, and two families of storm petrels. Formerly called Tubinares and still call ...
, along with
shearwater Shearwaters are medium-sized long-winged seabirds in the petrel family Procellariidae. They have a global marine distribution, but are most common in temperate and cold waters, and are pelagic outside the breeding season. Description These tub ...
s,
fulmar The fulmars are tubenosed seabirds of the family Procellariidae. The family consists of two extant species and two extinct fossil species from the Miocene. Fulmars superficially resemble gulls, but are readily distinguished by their flight on ...
s,
storm petrel Storm-petrel may refer to one of two bird families, both in the order Procellariiformes, once treated as the same family. The two families are: * Northern storm petrels (''Hydrobatidae'') are found in the Northern Hemisphere, although some speci ...
s, and
diving petrel The diving petrels form a genus, ''Pelecanoides'', of seabirds in the family Procellariidae. There are four very similar species of diving petrels, distinguished only by small differences in the coloration of their plumage, habitat, and bill co ...
s. They share certain identifying features. First, they have nasal passages that attach to the upper bill called
naricorns A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called tur ...
, although the nostrils of the albatross are on the sides of the bill. The bills of Procellariiformes are also unique in that they are split into between seven and nine horny plates. Finally, they produce a stomach oil made up of
wax ester A wax ester (WE) is an ester of a fatty acid and a fatty alcohol. Wax esters comprise the main components of three commercially important waxes: carnauba wax, candelilla wax, and beeswax.. Wax esters are formed by combining one fatty acid with on ...
s and
triglycerides A triglyceride (TG, triacylglycerol, TAG, or triacylglyceride) is an ester derived from glycerol and three fatty acids (from ''tri-'' and ''glyceride''). Triglycerides are the main constituents of body fat in humans and other vertebrates, as we ...
that is stored in the
proventriculus The proventriculus is part of the digestive system of birds.Encarta World English Dictionary orth American Edition(2007). ''Proventriculus''. Source: (accessed: December 18, 2007) An analogous organ exists in invertebrates and insects. Birds Th ...
. This is used against predators and as an energy rich food source for chicks and for the adults during their long flights. They also have a salt gland that is situated above the nasal passage and helps desalinate their bodies, due to the high amount of ocean water that they imbibe. It excretes a high saline solution from their nose.


Etymology

The waved albatross derives its name from the wave-like pattern of the feathers on adult birds.


Description

These are medium-sized albatrosses, measuring in length with a wingspan of .Howell (2012), p. 341. They range between in mass, with males averaging significantly heavier than females. They are distinctive for their yellowish-cream neck and head, which contrasts with their mostly brownish bodies. Even more distinctive is the very long, bright yellow bill, which looks disproportionately large in comparison to the relatively small head and long, slender neck. They also have chestnut brown upper parts and underparts, except for the breast, with fine barring, a little coarser on the rump. They have brown upper-wings, back, and tail, along with a whitish breast and underwings. Their
axilla The axilla (also, armpit, underarm or oxter) is the area on the human body directly under the shoulder joint. It includes the axillary space, an anatomical space within the shoulder girdle between the arm and the thoracic cage, bounded superior ...
ries are brown. Finally they have bluish feet. Juveniles are similar to adults except for more white on their head.BirdLife International (2008a) Chicks have brown fluffy feathers. The lifespan of this species may reach 40 to 45 years.


Range

The waved albatross breeds primarily on
Española Island Española Island (Spanish: ''Isla Española'') is part of the Galápagos Islands. The English named it ''Hood Island'' after Viscount Samuel Hood. It is located in the extreme southeast of the archipelago and is considered, along with Sant ...
in the Galápagos archipelago; however, there have been sightings of non-breeders and therefore possible small numbers, around 10 to 20, of breeders on Genovesa Island and Isla de la Plata.Clements, James (2007) During non-breeding season they will shift to the east and southeast to the continental shelf region off the coast of
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
and
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ' ...
. Sometimes they are seen in Colombia.


Behavior


Feeding

The primary food sources of the waved albatross are
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of ...
, squid, and
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group can ...
s, as well as smaller birds. But they have also been observed to scavenge for other food sources, including the regurgitated food of other birds. When foraging, the waved albatross finds places in the ocean where prey will be near this surface; this is the most effective way for the albatross to get its food. The waved albatrosses will forage away from the place where the chicks are nesting to get food for them.


Breeding

The nests are built on areas of lava with boulders and sparse vegetation, or thick brush. The courtship of the waved albatross is a very elusive and spectacular sight. It includes: rapid bill circling and bowing, beak clacking, and an upraised bill to make a whoo hoo sound. The eggs are laid between April and June and incubated for two months. When the eggs hatch, the chicks stay together in small nurseries while the parents go out to the sea for hunting. When the parents return, they may feed the chicks up to of oil. The young reach adult size by December and leave the colony by January. The partners remain mates until one of the partners dies."Galapagos Conservancy"


Flight

Waved albatross are spectacular flyers, perhaps even the most famous. They can fly for hours without stalling and they do this by
dynamic soaring Dynamic soaring is a flying technique used to gain energy by repeatedly crossing the boundary between air masses of different velocity. Such zones of wind gradient are generally found close to obstacles and close to the surface, so the technique i ...
. The wind speed near the surface of the sea is much lower than about in the air. The waved albatross uses this to its advantage by gliding at speed into the wind. As the waved albatross glides higher it loses most of its ground speed because it is gliding into a wind of a higher speed. However, its air speed does not fall, enabling it to glide continuously. Waved albatrosses do have difficulty in landing because of their high stalling speed, and in taking off because of their weight and wing span. To make it easier they sometimes take off from cliffs that are somewhat inland rather than beside the coast.


Conservation

The population of waved albatrosses on the Galápagos is protected by national park personnel, and the island is also categorized as a
World Heritage Site A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for h ...
. But limited range, bycatch by
longline fishing Longline fishing, or longlining, is a commercial fishing angling technique that uses a long ''main line'' with fish bait, baited fish hook, hooks attached at intervals via short branch lines called ''snoods'' or ''gangions''.< ...
, disturbance via tourism, disease, and the effects of
illegal fishing Illegal, unreported and unregulated fishing (IUU) is an issue around the world. Fishing industry observers believe IUU occurs in most fisheries, and accounts for up to 30% of total catches in some important fisheries. Illegal fishing takes ...
in the nearby waters place them in considerable jeopardy. Longline fishing in particular seems to be having a severe impact on the species, the conservation status of which was upgraded from
near threatened A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify f ...
to vulnerable by the IUCN in 2000. Despite there still being some 34,700 adult birds in 2001, their numbers have apparently started to decrease at an unknown rate more recently, probably due to longline fishing which also upsets the
sex ratio The sex ratio (or gender ratio) is usually defined as the ratio of males to females in a population. As explained by Fisher's principle, for evolutionary reasons this is typically about 1:1 in species which reproduce sexually. Many species d ...
(males being killed more frequently). As the current situation makes the population highly vulnerable to a catastrophic collapse to
extinction Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
, it was uplisted to critically endangered status in the 2007
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
. The population of 34,700 adult birds was based on a 2001 estimate; however in 1970 and 1971 there were an estimated 24,000, and 1994 saw between 31,200 and 36,400. This species has an occurrence range of , and a breeding range of .


Gallery

File:Phoebastria irrorata -Espanola Island, Galapagos Islands, Ecuador -flying-8.jpg, In flight File:Waved-chick.jpg, Unfledged chick in January File:Waved-albatross-egg.jpg, Abandoned egg File:DiomedeaIrrorataSmit.jpg, Illustration by
Joseph Smit Joseph Smit (18 July 1836 – 4 November 1929) was a Dutch zoological illustrator. L.B. Holthuis, Leiden, (1958, 1995) ''Rijksmuseum van Natuurlijke Historie, 1820 - 1958''. page 47reprint manuscript, PDF Background Smit was born in Lisse. H ...
, 1896


Footnotes


References

* * * *
Darwin Foundation – Albatross
* * * * *


External links


Species factsheet
- BirdLife International {{Taxonbar, from=Q735296 waved albatross Birds of Ecuador Birds of Peru Galápagos Islands coastal fauna waved albatross waved albatross