''Wargosuchus'' (meaning "

warg In the philologist and fantasy author J. R. R. Tolkien's Middle-earth fiction, a warg is a particularly large and evil kind of wolf that could be ridden by orcs. He derived the name and characteristics of his wargs by combining meanings and myth ...

crocodile Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term crocodile is sometimes used even more loosely to inclu ...

") is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

baurusuchid Baurusuchidae is a Gondwanan Family (biology), family of mesoeucrocodylians that lived during the Late Cretaceous. It is a group of terrestrial hypercarnivorous crocodilians from South America (Argentina and Brazil) and possibly Pakistan. Baurusu ...

mesoeucrocodylia Mesoeucrocodylia is the clade that includes Eusuchia and crocodyliforms formerly placed in the paraphyletic group Mesosuchia. The group appeared during the Early Jurassic, and continues to the present day. Diagnosis It was long known that M ...

n from the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

. It is known from a fragmentary skull from the

Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. The ...

-age

Bajo de la Carpa Formation The Bajo de la Carpa Formation is a geologic formation of the Neuquén Basin that crops out in northern Patagonia, in the provinces of Río Negro and Neuquén, Argentina. It is the oldest of two formations belonging to the Río Colorado Subgroup ...

of the

Neuquén Group The Neuquén Group is a group of geologic formations found in Argentina. Rocks in the Neuquén Group fall within the Cenomanian to early Campanian stages of the Late Cretaceous Period. It overlies the older Lohan Cura Formation and is itself over ...

, found in the vicinity of

Neuquén Neuquén (; arn, Nehuenken) is the capital city of the Argentine province of Neuquén and of the Confluencia Department, located in the east of the province. It occupies a strip of land west of the confluence of the Limay and Neuquén rivers w ...

Neuquén Province Neuquén () is a province of Argentina, located in the west of the country, at the northern end of Patagonia. It borders Mendoza Province to the north, Rio Negro Province to the southeast, and Chile to the west. It also meets La Pampa Province a ...

, and was described by Agustín Martinelli and Diego Pais in 2008. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

, and so far the only species, is ''Wargosuchus australis''. ''Wargosuchus'' is based on MOZ-PV 6134, a partial right

premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has b ...

and

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...

, and partial skull roof. Martinelli and Pais distinguished ''Wargosuchus'' from other mesoeucrocodylians by skull details, such as a deep groove on the midline of the

frontal bone The frontal bone is a bone in the human skull. The bone consists of two portions.''Gray's Anatomy'' (1918) These are the vertically oriented squamous part, and the horizontally oriented orbital part, making up the bony part of the forehead, par ...

s, a large depression for the

olfactory bulb The olfactory bulb (Latin: ''bulbus olfactorius'') is a grey matter, neural structure of the vertebrate forebrain involved in olfaction, the sense of odor, smell. It sends olfactory information to be further processed in the amygdala, the orbitof ...

s, and enlargement of the last tooth of the premaxilla followed by a deep pit for the following tooth of the lower jaw. The animal had a robust skull. It shared its setting with four other taxa of mesoeucrocodylians: common ''

Notosuchus ''Notosuchus'' (; 'southern crocodile') is an extinct genus of South American notosuchian crocodylomorph. It was terrestrial, living approximately 85 million years ago in the Santonian stage of the Late Cretaceous. Description ''Notosuchus'' wa ...

'', and rare ''

Cynodontosuchus ''Cynodontosuchus'' is an extinct genus of baurusuchid mesoeucrocodylian. Fossils have been found from Argentina of Late Cretaceous age from the Bajo de la Carpa Formation (dating back to the Santonian), the Pichi Picun Leufu Formation (dating ...

'', ''

Lomasuchus ''Lomasuchus'' is an extinct genus of peirosaurid notosuchian known from the Late Cretaceous (Santonian stage) of Neuquén Province, western central Argentina. It contains a single species, ''Lomasuchus palpebrosus''. Gasparini, Z., Chiappe, L. M ...

'', and ''

Gasparinisuchus ''Gasparinisuchus'' is an extinct genus of peirosaurid notosuchian known from the Late Cretaceous (Santonian to early Campanian stage) of Neuquén Province, Neuquén and Mendoza Provinces, western central Argentina. It contains a single species, ...

''.

References

{{Taxonbar, from=Q2783787 Late Cretaceous crocodylomorphs of South America Terrestrial crocodylomorphs Notosuchians Prehistoric pseudosuchian genera