''Waimanu'' is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of early
penguin
Penguins (order (biology), order List of Sphenisciformes by population, Sphenisciformes , family (biology), family Spheniscidae ) are a group of Water bird, aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: on ...
which lived during the
Paleocene
The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
, soon after the
Cretaceous–Paleogene extinction event
The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...
, around 62–60 million years ago. It was about the size of an
emperor penguin
The emperor penguin (''Aptenodytes forsteri'') is the tallest and heaviest of all living penguin species and is endemic to Antarctica. The male and female are similar in plumage and size, reaching in length and weighing from . Feathers of th ...
(1 metre). It is one of the most important bird fossils for understanding the origin and evolution of birds because of the time period it comes from, and the position of penguins near the base of the bird family tree.
''Waimanu'' was a very early member of the
Sphenisciformes
Penguins ( order Sphenisciformes , family Spheniscidae ) are a group of aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: only one species, the Galápagos penguin, is found north of the Equator. Highly adapt ...
, the order that includes modern penguins. However, although it was probably flightless like all modern penguins, with wings specialized for wing-propelled diving, its wing bones do not yet show the extreme specializations modern penguins have for an aquatic lifestyle.
[ It may have resembled a flightless ]loon
Loons (North American English) or divers (British / Irish English) are a group of aquatic birds found in much of North America and northern Eurasia. All living species of loons are members of the genus ''Gavia'', family Gaviidae and order Gavi ...
or diver in body shape, and possibly the great auk
The great auk (''Pinguinus impennis'') is a species of flightless alcid that became extinct in the mid-19th century. It was the only modern species in the genus ''Pinguinus''. It is not closely related to the birds now known as penguins, wh ...
in its manner of locomotion.
Discovery
''Waimanu'' was discovered in the Basal Waipara Greensand
The Waipara Greensand is a geological rock unit found in Canterbury, New Zealand. It dates from just after the Cretaceous–Paleogene extinction event, the Thanetian and Selandian, around 61–62 million years ago in the early Palaeocene. It is w ...
near the Waipara River, in Canterbury
Canterbury (, ) is a City status in the United Kingdom, cathedral city and UNESCO World Heritage Site, situated in the heart of the City of Canterbury local government district of Kent, England. It lies on the River Stour, Kent, River Stour.
...
, New Zealand
New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
, in 1980.
The holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
is a partial skeleton comprising almost complete right tibiotarsus, proximal half of right fibula, right tarsometatarsus, right pelvis, and synsacrum (with last thoracic vertebra attached to the synsacrum), four caudal vertebrae. It is held in the Canterbury Museum in Christchurch, New Zealand.
Etymology
The genus name ''Waimanu'' comes from the Māori
Māori or Maori can refer to:
Relating to the Māori people
* Māori people of New Zealand, or members of that group
* Māori language, the language of the Māori people of New Zealand
* Māori culture
* Cook Islanders, the Māori people of the C ...
''wai'' for "water" and ''manu'' for "bird". One species is known, ''W. manneringi'', named for Al Mannering who found and collected the holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen A second species, ''Waimanu tuatahi'', was moved to ''Muriwaimanu
''Muriwaimanu'' is an extinct genus of early penguin.
One species is known, ''Muriwaimanu tuatahi'', which was originally referred to ''Waimanu tuatahi'' in 2006 by Slack ''et al.''.Slack, K.E., Jones, C.M., Ando, T., Harrison G.L., Fordyce R.E. ...
'' in 2018.
Significance
The discovery of ''Waimanu'' provided evidence for the debate about whether the radiation of modern birds, Neoaves
Neoaves is a clade that consists of all modern birds (Neornithes or Aves) with the exception of Paleognathae (ratites and kin) and Galloanserae (ducks, chickens and kin). Almost 95% of the roughly 10,000 known species of extant birds belong to ...
, took place before the extinction of the non-avian dinosaurs
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
or extremely rapidly immediately after. DNA studies, combined with the fossil evidence, seem to indicate the latter.
References
External links
Images of ''Waimanu'' specimens and a phylogram of early birds.
Skull of world's oldest penguin found
{{Taxonbar, from=Q138638
Prehistoric bird genera
Sphenisciformes
Paleocene birds
Extinct penguins
Extinct birds of New Zealand
Fossil taxa described in 2006