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''Voyria'', commonly known as ghostplants, is a genus of 20 species of
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
s, belonging to the family
Gentianaceae Gentianaceae is a family of flowering plants of 103 genera and about 1600 species. Etymology The family takes its name from the genus '' Gentiana'', named after the Illyrian king Gentius. Distribution Distribution is cosmopolitan. Characteri ...
. They are mostly native to warm temperate and tropical regions of the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
,
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, except for '' V. primuloides'', which is found in West and Central Africa. ''V. parasitica'' reaches as far north as the
Everglades The Everglades is a natural region of tropical climate, tropical wetlands in the southern portion of the U.S. state of Florida, comprising the southern half of a large drainage basin within the Neotropical realm. The system begins near Orland ...
in
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
. Unlike most plants, they do not contain
chlorophyll Chlorophyll (also chlorophyl) is any of several related green pigments found in cyanobacteria and in the chloroplasts of algae and plants. Its name is derived from the Greek words , ("pale green") and , ("leaf"). Chlorophyll allow plants to a ...
; they are
myco-heterotrophs Myco-heterotrophy (from Greek μύκης , "fungus", ἕτερος ', "another", "different" and τροφή ', "nutrition") is a symbiotic relationship between certain kinds of plants and fungi, in which the plant gets all or part of its food fr ...
, getting their food through
parasitism Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted structurally to this way of lif ...
upon
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
rather than
photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored i ...
. Their roots are thick and densely clustered, forming a "birds nest" that house their fungus host. Depending on the species, the flowers can be single or held in corymbs of many individual flowers. The stems are usually pallid, with reduced scale-like foliage. The flowers can vary in colour, with white or yellow predominating, and blue and pink also occurring. Like all myco-heterotrophs, they are capable of living in very dark conditions, such as the floor of deep
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s, because they no longer derive energy from sunlight. ''Voyria'' is subdivided into two subgenera, ''Voyria'' and ''Leiphaimos''. The latter, which contains a majority of the species, is characterized by highly reduced features, lacking both
stoma In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bor ...
tes and a continuous
vascular The blood vessels are the components of the circulatory system that transport blood throughout the human body. These vessels transport blood cells, nutrients, and oxygen to the tissues of the body. They also take waste and carbon dioxide away f ...
cylinder. The lack of
chloroplast genes A chloroplast () is a type of membrane-bound organelle known as a plastid that conducts photosynthesis mostly in plant and algal cells. The photosynthetic pigment chlorophyll captures the energy from sunlight, converts it, and stores it in ...
has caused some difficulty in better understanding its relationships within the family. Seeds are dispersed by wind.


Species list

*''
Voyria acuminata ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South ...
'' Benth. -
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
*''
Voyria alvesiana ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South ...
'' E.F.Guim., T.S.Mendes & N.G.Silva -
Pará Pará is a Federative units of Brazil, state of Brazil, located in northern Brazil and traversed by the lower Amazon River. It borders the Brazilian states of Amapá, Maranhão, Tocantins (state), Tocantins, Mato Grosso, Amazonas (Brazilian state) ...
*''
Voyria aphylla ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South ...
'' (Jacq.) Pers. -
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
,
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
to
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
*'' Voyria aurantiaca'' Splitg. -
Lesser Antilles The Lesser Antilles ( es, link=no, Antillas Menores; french: link=no, Petites Antilles; pap, Antias Menor; nl, Kleine Antillen) are a group of islands in the Caribbean Sea. Most of them are part of a long, partially volcanic island arc betwe ...
,
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
to
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
*''
Voyria caerulea ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South A ...
'' Aubl. -
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
,
the Guianas The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France * ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
*''
Voyria chionea ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South A ...
'' Benth. -
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
,
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
*''
Voyria clavata ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South A ...
'' Splitg. -
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
,
the Guianas The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France * ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
*''
Voyria corymbosa ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South A ...
'' Splitg. -
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
,
the Guianas The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France * ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
*''
Voyria crucitasensis ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South A ...
'' Y.Guillén & G.Vargas - N
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
*''
Voyria flavescens ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South A ...
'' Griseb. -
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
to
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
,
Trinidad Trinidad is the larger and more populous of the two major islands of Trinidad and Tobago. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is often referred to as the southernmos ...
*'' Voyria kupperi'' (Suess.) Ruyters & Maas -
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
,
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
*''
Voyria obconica ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South A ...
'' Progel -
Atlantic Forest The Atlantic Forest ( pt, Mata Atlântica) is a South American forest that extends along the Atlantic coast of Brazil from Rio Grande do Norte state in the northeast to Rio Grande do Sul state in the south and inland as far as Paraguay and th ...
of
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
*''
Voyria parasitica ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South ...
'' (Schltdl. & Cham.) Ruyters & Maas -
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
,
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
, S
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
*''
Voyria primuloides ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South ...
'' Baker -
Sierra Leone Sierra Leone,)]. officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered by Liberia to the southeast and Guinea surrounds the northern half of the nation. Covering a total area of , Sierra ...
,
Liberia Liberia (), officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to its north, Ivory Coast to its east, and the Atlantic Ocean ...
,
Ivory Coast Ivory Coast, also known as Côte d'Ivoire, officially the Republic of Côte d'Ivoire, is a country on the southern coast of West Africa. Its capital is Yamoussoukro, in the centre of the country, while its largest city and economic centre is ...
,
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
,
Equatorial Guinea Equatorial Guinea ( es, Guinea Ecuatorial; french: Guinée équatoriale; pt, Guiné Equatorial), officially the Republic of Equatorial Guinea ( es, link=no, República de Guinea Ecuatorial, french: link=no, République de Guinée équatoria ...
,
Gabon Gabon (; ; snq, Ngabu), officially the Gabonese Republic (french: République gabonaise), is a country on the west coast of Central Africa. Located on the equator, it is bordered by Equatorial Guinea to the northwest, Cameroon to the north ...
,
Congo Republic The Republic of the Congo (french: République du Congo, ln, Republíki ya Kongó), also known as Congo-Brazzaville, the Congo Republic or simply either Congo or the Congo, is a country located in the western coast of Central Africa to the w ...
*''
Voyria pulcherrima ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South A ...
'' (Standl.) L.O.Williams -
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
*'' Voyria rosea'' Aubl. -
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
and
the Guianas The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France * ...
*''
Voyria spruceana ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South A ...
'' Benth. -
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
to
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
*'' Voyria tenella'' Guilding ex Hook. -
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
,
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
to
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
*''
Voyria tenuiflora ''Voyria'', commonly known as ghostplants, is a genus of 20 species of herbaceous perennial plants, belonging to the family Gentianaceae. They are mostly native to warm temperate and tropical regions of the Caribbean, Central America and South A ...
'' Griseb. -
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
,
the Guianas The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France * ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
*'' Voyria truncata'' (Standl.) Standl. & Steyerm. -
Chiapas Chiapas (; Tzotzil language, Tzotzil and Tzeltal language, Tzeltal: ''Chyapas'' ), officially the Free and Sovereign State of Chiapas ( es, Estado Libre y Soberano de Chiapas), is one of the states that make up the Political divisions of Mexico, ...
to
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...


References

{{Taxonbar, from=Q6799774 Gentianaceae Parasitic plants Flora of North America Gentianaceae genera Flora of South America Flora of Africa Flora of the Caribbean Flora of Mexico Flora of Central America