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The Visayan hornbill (''Penelopides panini'') is a
hornbill Hornbills (Bucerotidae) are a family (biology), family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a Casque (an ...
found in
rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...
s on the islands of
Panay Panay is the sixth-largest and fourth-most populous island in the Philippines, with a total land area of and has a total population of 4,542,926 as of 2020 census. Panay comprises 4.4 percent of the entire population of the country. The City o ...
,
Negros Negros is the fourth largest and third most populous island in the Philippines, with a total land area of . Negros is one of the many islands of the Visayas, in the central part of the country. The predominant inhabitants of the island region a ...
,
Masbate Masbate, officially the Province of Masbate ( Masbateño: ''Probinsya san Masbate''; tl, Lalawigan ng Masbate), is an island province in the Philippines located near the midsection of the nation's archipelago. Its provincial capital is Masbate C ...
, and
Guimaras Guimaras , officially the Province of Guimaras ( hil, Kapuoran sang Guimaras; tl, Lalawigan ng Guimaras), is an island province in the Philippines located in the Western Visayas region. Its capital is Jordan while its largest local government ...
, and formerly Ticao, in the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
. It formerly included all other Philippine
tarictic hornbill ''Penelopides'' is the genus of relatively small, primarily frugivorous hornbills restricted to forested areas of the Philippines. Their common name, tarictic hornbills, is an onomatopoetic reference to the main call of several of them. They have ...
s as
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
, in which case the
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
of the 'combined species' was shortened to tarictic hornbill.


Taxonomy

The Visayan hornbill was described by the French polymath
Georges-Louis Leclerc, Comte de Buffon Georges-Louis Leclerc, Comte de Buffon (; 7 September 1707 – 16 April 1788) was a French naturalist, mathematician, cosmologist, and encyclopédiste. His works influenced the next two generations of naturalists, including two prominent Fr ...
in 1780 in his ''Histoire Naturelle des Oiseaux''. The bird was also illustrated in a hand-coloured plate engraved by
François-Nicolas Martinet François-Nicolas Martinet (1731 - 1800) was a French engineer, engraver and naturalist. Martinet engraved the plates for numerous works on natural history, especially ornithology. Notable in particular are those for ''l'Ornithologia, sive Synop ...
in the ''Planches Enluminées D'Histoire Naturelle'' which was produced under the supervision of
Edme-Louis Daubenton Edme-Louis Daubenton (12 August 1730 – 12 December 1785) was a French naturalist. Daubenton was the cousin of another French naturalist, Louis Jean-Marie Daubenton. Georges-Louis Leclerc, the Comte de Buffon engaged Edme-Louis Daubenton to su ...
to accompany Buffon's text. Neither the plate caption nor Buffon's description included a scientific name but in 1783 the Dutch naturalist
Pieter Boddaert Pieter Boddaert (1730 – 6 May 1795) was a Dutch physician and natural history, naturalist. Early life, family and education Boddaert was the son of a Middelburg jurist and poet by the same name (1694–1760). The younger Pieter obtained his M.D ...
coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Buceros panini'' in his catalogue of the ''Planches Enluminées''. The type locality is the island of
Panay Panay is the sixth-largest and fourth-most populous island in the Philippines, with a total land area of and has a total population of 4,542,926 as of 2020 census. Panay comprises 4.4 percent of the entire population of the country. The City o ...
in the Philippines. The Visayan hornbill is now placed in the genus ''Penelopides'' that was introduced in 1849 by the German naturalist
Ludwig Reichenbach Heinrich Gottlieb Ludwig Reichenbach (8 January 1793 – 17 March 1879) was a German botanist and ornithologist. It was he who first requested Leopold Blaschka to make a set of glass marine invertebrate models for scientific education and museu ...
in a plate of the hornbills. The origin of the generic name is uncertain but it may be a combination of the Latin ''pene'' meaning "almost" or "nearly", the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''lophos'' meaning "crest" and ''-oideēs'' "resembling". The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''panini'' is a Latinized form of "Panay". The common English name refers to the
Visayan Islands The Visayas ( ), or the Visayan Islands (Visayan: ''Kabisay-an'', ; tl, Kabisayaan ), are one of the three principal geographical divisions of the Philippines, along with Luzon and Mindanao. Located in the central part of the archipelago, it ...
. These are located in the central part of the Philippines and include the island of Panay. Two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognized: * ''P. p. panini'' – (
Boddaert Pieter Boddaert (1730 – 6 May 1795) was a Dutch physician and naturalist. Early life, family and education Boddaert was the son of a Middelburg jurist and poet by the same name (1694–1760). The younger Pieter obtained his M.D. at the Univers ...
, 1783)
: Visayan tarictic hornbill,
nominate Nomination is part of the process of selecting a candidate for either election to a public office, or the bestowing of an honor or award. A collection of nominees narrowed from the full list of candidates is a short list. Political office In the ...
, found on
Panay Panay is the sixth-largest and fourth-most populous island in the Philippines, with a total land area of and has a total population of 4,542,926 as of 2020 census. Panay comprises 4.4 percent of the entire population of the country. The City o ...
,
Negros Negros is the fourth largest and third most populous island in the Philippines, with a total land area of . Negros is one of the many islands of the Visayas, in the central part of the country. The predominant inhabitants of the island region a ...
,
Masbate Masbate, officially the Province of Masbate ( Masbateño: ''Probinsya san Masbate''; tl, Lalawigan ng Masbate), is an island province in the Philippines located near the midsection of the nation's archipelago. Its provincial capital is Masbate C ...
and
Guimaras Guimaras , officially the Province of Guimaras ( hil, Kapuoran sang Guimaras; tl, Lalawigan ng Guimaras), is an island province in the Philippines located in the Western Visayas region. Its capital is Jordan while its largest local government ...
* ''P. p. ticaensis'' – Hachisuka, 1930: Ticao tarictic hornbill, found on
Ticao Island Ticao Island is an island with a total land area of . It is one of the three major islands of Masbate province in the Philippines. It is separated from the Bicol Peninsula by the Ticao Pass. The other two major islands are Masbate Island () and ...
(likely
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
)


Description

The adults show
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
. The male has a creamy-white head and neck, a white upper chest, a reddish-brown lower chest and uppertail-
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are s ...
, and a creamy-white buff tail with a broad black tip. The bill and casque are blackish; the former with yellowish ridges. The bare ocular skin is pinkish-white. The tail and bill of the female resemble that of the male, but otherwise the
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
of the female is black, and the ocular skin is blue.


Diet and behavior

Visayan hornbills live in groups and frequent the canopy of rainforests. These birds are noisy and emit an incessant sound that sounds like ''ta-rik-tik'', hence the name. Despite their noise they are difficult to find, being well camouflaged by the dense
foliage A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...
. The principal food of Visayan hornbill is fruit. It also eats
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s,
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s,
ant Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22 ...
s and
earthworm An earthworm is a terrestrial invertebrate that belongs to the phylum Annelida. They exhibit a tube-within-a-tube body plan; they are externally segmented with corresponding internal segmentation; and they usually have setae on all segments. Th ...
s (rarely).


Conservation

This is a highly endangered species. The total population is estimated at 1800 individuals. There has been a heavy decline in population due to
hunting Hunting is the human activity, human practice of seeking, pursuing, capturing, or killing wildlife or feral animals. The most common reasons for humans to hunt are to harvest food (i.e. meat) and useful animal products (fur/hide (skin), hide, ...
and loss of habitat caused by
deforestation Deforestation or forest clearance is the removal of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. The most concentrated d ...
. The subspecies ''ticaensis'' was described as "abundant" in 1905, but almost the entire forest on the island was replaced by plantations and settlements in the 20th century. The last time the Ticao tarictic was seen was in 1971, and it is now likely to be
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
. If confirmed, this is the first
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
of hornbill to go extinct in recorded history; many other taxa in the family are now at risk.


Captivity

This species has just been imported from Panay in the Philippines by
Chester Zoo Chester Zoo is a zoo at Upton-by-Chester, Cheshire, England. Chester Zoo was opened in 1931 by George Mottershead and his family. It is one of the UK's largest zoos at . The zoo has a total land holding of approximately . Chester Zoo is operate ...
, England. There are two pairs at Chester, and two pairs at
Avifauna Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight s ...
in the Netherlands, amongst other collections. In the past, the
Los Angeles Zoo The Los Angeles Zoo and Botanical Gardens is a zoo founded in 1966 and located in Los Angeles, California. The city of Los Angeles owns the entire zoo, its land and facilities, and the animals. Animal care, grounds maintenance, construction, ed ...
has bred this species, but it is not known whether these birds were pure ''Penelopides panini panini'', so it may not be the first captive breeding of this species; that title may go to a breeding centre on Panay, where Chester's birds came from. The Chester zoo has bred this species.


References

*


External links

* * * * * {{Taxonbar, from=Q647138
Visayan hornbill The Visayan hornbill (''Penelopides panini'') is a hornbill found in rainforests on the islands of Panay Island, Panay, Negros Island, Negros, Masbate, and Guimaras Island, Guimaras, and formerly Ticao Island, Ticao, in the Philippines. It former ...
Fauna of the Visayas Birds of Panay Birds of Negros Island Fauna of Guimaras Fauna of Masbate
Visayan hornbill The Visayan hornbill (''Penelopides panini'') is a hornbill found in rainforests on the islands of Panay Island, Panay, Negros Island, Negros, Masbate, and Guimaras Island, Guimaras, and formerly Ticao Island, Ticao, in the Philippines. It former ...