Vipera Ursinii
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:''Common names: meadow viper, Ursini's viper, Mallow D, Ludwig D, Nilson G (2003). ''True Vipers: Natural History and Toxinology of Old World Vipers''. Malabar, Florida: Krieger Publishing Company. . meadow adder,Brown, John Haynes (1973). ''Toxicology and Pharmacology of Venoms from Poisonous Snakes''. Springfield, Illinois: Charles C. Thomas. . (
more More or Mores may refer to: Computing * MORE (application), outline software for Mac OS * more (command), a shell command * MORE protocol, a routing protocol * Missouri Research and Education Network Music Albums * ''More!'' (album), by Booka ...
).'' ''Vipera ursinii'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
venomous snake Venomous snakes are Species (biology), species of the Suborder (biology), suborder Snake, Serpentes that are capable of producing Snake venom, venom, which they use for killing prey, for defense, and to assist with digestion of their prey. The v ...
in the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Viperinae The Viperinae, or viperines, are a subfamily of vipers endemic to Europe, Asia and Africa. They are distinguished by their lack of the heat-sensing pit organs that characterize their sister group, the subfamily Crotalinae. Currently, 13 genera a ...
of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Viperidae The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs tha ...
. It is a very rare species, which is in danger of
extinction Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
. This species is commonly called the meadow viper. It is found in
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
, and
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
as well as much of
eastern Europe Eastern Europe is a subregion of the Europe, European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural, and socio-economic connotations. The vast majority of the region is covered by Russ ...
. Several subspecies are recognized. Beyond the highly threatened European population, poorly known populations exist as far to the east as
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
and northwestern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
.


Etymology

The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
or epithet, ''ursinii'', is in honor of Italian naturalist
Antonio Orsini Antonio is a masculine given name of Etruscan origin deriving from the root name Antonius. It is a common name among Romance language-speaking populations as well as the Balkans and Lusophone Africa. It has been among the top 400 most popular male ...
(1788–1870).


Description

Adults of ''V. ursini'' average in total length (including tail), although specimens of in total length have been reported. Females are larger than males. Although sometimes confused with '' V. aspis'' or '' V. berus'', it differs from them in the following characters. The smallest viper in Europe, its body is thick, its head narrow, and its appearance rough. The snout is not upturned. There are always several large scales or plates on the top of the head. The prominently keeled dorsal scales are in only 19 rows, and often dark skin shows between them. It is gray, tan, or yellowish with a dark undulating dorsal stripe, which is edged with black.


Common names

Meadow viper, Ursini's viper, meadow adder, Orsini's viper, field viper, Steward JW (1971). ''The Snakes of Europe''. Cranbury, New Jersey: Associated University Press (Fairleigh Dickinson University Press). . field adder. Hellmich W (1962). ''Reptiles and Amphibians of Europe''. London: Blandford Press. ''Translated from'' Winter C (1956). ''Die Lurche und Kriechtiere Europas''. Heidelberg, Germany: Universitatsverlag, gegr. 1822, GmbH. Although the following subspecies are currently invalid according to the taxonomy used here, their common names may still be encountered: * ''V. u. ursinii'' – Italian meadow viper. * ''V. u. macrops'' – karst viper, karst adder. * ''V. u. rakosiensis'' – Danubian meadow viper. * ''V. renardi'' – steppe viper, steppe adder, Renard's viper. * ''V. u. moldavica'' – Moldavian meadow viper.


Geographic range

Southeastern
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
, eastern
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
(extinct),
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...
, central
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
,
Serbia Serbia (, ; Serbian language, Serbian: , , ), officially the Republic of Serbia (Serbian language, Serbian: , , ), is a landlocked country in Southeast Europe, Southeastern and Central Europe, situated at the crossroads of the Pannonian Bas ...
,
Montenegro ) , image_map = Europe-Montenegro.svg , map_caption = , image_map2 = , capital = Podgorica , coordinates = , largest_city = capital , official_languages = M ...
,
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...
,
Bosnia and Herzegovina Bosnia and Herzegovina ( sh, / , ), abbreviated BiH () or B&H, sometimes called Bosnia–Herzegovina and often known informally as Bosnia, is a country at the crossroads of south and southeast Europe, located in the Balkans. Bosnia and H ...
, northern and northeastern
Republic of Kosovo Kosovo ( sq, Kosova or ; sr-Cyrl, Косово ), officially the Republic of Kosovo ( sq, Republika e Kosovës, links=no; sr, Република Косово, Republika Kosovo, links=no), is a international recognition of Kosovo, partiall ...
,
North Macedonia North Macedonia, ; sq, Maqedonia e Veriut, (Macedonia before February 2019), officially the Republic of North Macedonia,, is a country in Southeast Europe. It gained independence in 1991 as one of the successor states of Socialist Feder ...
,
Albania Albania ( ; sq, Shqipëri or ), or , also or . officially the Republic of Albania ( sq, Republika e Shqipërisë), is a country in Southeastern Europe. It is located on the Adriatic and Ionian Seas within the Mediterranean Sea and shares ...
,
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...
, northern
Bulgaria Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Macedon ...
,
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
,
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
, northwestern
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
,
Armenia Armenia (), , group=pron officially the Republic of Armenia,, is a landlocked country in the Armenian Highlands of Western Asia.The UNbr>classification of world regions places Armenia in Western Asia; the CIA World Factbook , , and ''Ox ...
,
Azerbaijan Azerbaijan (, ; az, Azərbaycan ), officially the Republic of Azerbaijan, , also sometimes officially called the Azerbaijan Republic is a transcontinental country located at the boundary of Eastern Europe and Western Asia. It is a part of th ...
, Georgia,
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
and across
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
,
Kyrgyzstan Kyrgyzstan,, pronounced or the Kyrgyz Republic, is a landlocked country in Central Asia. Kyrgyzstan is bordered by Kazakhstan to the north, Uzbekistan to the west, Tajikistan to the south, and the People's Republic of China to the east. ...
and eastern
Uzbekistan Uzbekistan (, ; uz, Ozbekiston, italic=yes / , ; russian: Узбекистан), officially the Republic of Uzbekistan ( uz, Ozbekiston Respublikasi, italic=yes / ; russian: Республика Узбекистан), is a doubly landlocked cou ...
steppes to
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
(
Xinjiang Xinjiang, SASM/GNC: ''Xinjang''; zh, c=, p=Xīnjiāng; formerly romanized as Sinkiang (, ), officially the Xinjiang Uygur Autonomous Region (XUAR), is an autonomous region of the People's Republic of China (PRC), located in the northwest ...
). ''Vipera ursinii rakosiensis'' is native to
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...
Leírás
Fertő-Hanság Nemzeti Park (Fertő-Hansag National Park of Hungary)
although the taxonomic status of this subspecies is disputed (see section "Taxonomy") The type locality is " ...''monti dell'Abruzzo prossimi alla provincia d'Ascoli''... " (...mountains of
Abruzzo Abruzzo (, , ; nap, label=Neapolitan language, Abruzzese Neapolitan, Abbrùzze , ''Abbrìzze'' or ''Abbrèzze'' ; nap, label=Sabino dialect, Aquilano, Abbrùzzu; #History, historically Abruzzi) is a Regions of Italy, region of Southern Italy wi ...
near the
Province of Ascoli Piceno The province of Ascoli Piceno ( it, Provincia di Ascoli Piceno) is a province in the Marche region of Italy. Its capital is the city of Ascoli Piceno, and the province is bordered by the Adriatic Sea to the east, the Province of Fermo to the north ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
...).


Conservation status

The species ''Vipera ursinii'' is considered to be a
Vulnerable species A vulnerable species is a species which has been Conservation status, categorized by the International Union for Conservation of Nature as being threatened species, threatened with extinction unless the circumstances that are threatened species, ...
on the
IUCN Red List of Threatened Species The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biologi ...
, due to
habitat destruction Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
caused by changes in agricultural practices and climate change in mountain areas, and to collection for the pet trade. In addition, this species is listed on
CITES CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
Appendix I, which prohibits commercial international trade,''Vipera ursinii''
a
CITES
an
United Nations Environment Programme / World Conservation Monitoring Centre
Accessed 8 October 2006.
and is a strictly protected species (Appendix II) under the Berne Convention.Convention on the Conservation of European Wildlife and Natural Habitats, Appendix II
a
Council of Europe
Accessed 9 October 2006.
''V. ursinii'' is the most threatened snake in Europe. At least 12 human activities are threatening these animals: 1. Grazing 2. Mowing 3. Fire 4. Agriculture 5. Roads 6. Constructions 7. Leisure Activities 8. Afforestation 9. Cynegetic species management 10. Persecution 11. Illegal collection 12. Littering


Taxonomy

There is high genetic diversity within samples of ''Vipera ursinii'' and several species may be involved. At least six subspecies may be encountered in modern literature: * ''Vipera ursinii ursinii'' ( Bonaparte, 1835) * ''Vipera ursinii eriwanensis'' ( A.F. Reuss, 1933) * ''Vipera ursinii graeca'' Nilson & Andrén, 1988 * ''Vipera ursinii macrops'' Méhelÿ, 1911 * ''Vipera ursinii moldavica'' Nilson, Andrén & Joger, 1993 * ''
Vipera ursinii rakosiensis The Hungarian meadow viper (''Vipera ursinii rakosiensis''; Hungarian language, Hungarian name: ''rákosi vipera''), also called the Danubian meadow viperSteward JW (1971). ''The Snakes of Europe''. Cranbury, New Jersey: Associated University P ...
'' Méhely, 1893 * ''Vipera ursinii renardi''
Christoph Christoph is a male given name and surname. It is a German variant of Christopher. Notable people with the given name Christoph * Christoph Bach (1613–1661), German musician * Christoph Büchel (born 1966), Swiss artist * Christoph Dientzenho ...
, 1861
Golay et al. (1993) recognize the first four, while Mallow et al. (2003) recognize five and list ''V. eriwanensis'' and ''V. renardi'' as valid species. However, McDiarmid et al. (1999), and thus ITIS, feel that more definitive data is necessary before any subspecies can be recognized.


Notes


References


Further reading

* Golay P, Smith HM, Broadley DG, Dixon JR, McCarthy CJ, Rage J-C, Schätti B, Toriba M (1993). ''Endoglyphs and Other Major Venomous Snakes of the World: A Checklist''. Geneva: Azemiops. * Latifi M (1991). ''The Snakes of Iran''. Oxford, Ohio: Society for the Study of Amphibians and Reptiles. . (''Vipera ursinii'', p. 133.)


External links

*
Meadow viper (''Vipera ursinii '')
a
ARKive
Accessed 5 October 2006.

a
Amphibians and Reptiles of Europe
Accessed 9 October 2006.

at ttps://web.archive.org/web/20070808155828/http://www.reptilia-amphibia.net/ Reptiles & Amphibians of France Accessed 30 October 2006. *
''Vipera eriwanensis''
a

Accessed 30 March 2007. {{Authority control ursinii Reptiles of Europe Reptiles of Central Asia Reptiles of Azerbaijan Reptiles described in 1835 Taxa named by Charles Lucien Bonaparte