:''Common names: Central Italian asp.Mehrtens JM. 1987. ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. .'' ''Vipera aspis francisciredi'' is a

venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

of the

European asp ''Vipera aspis'' is a Viperinae, viper species found in southwestern Europe. Its common names include asp, asp viper,Mallow D, Ludwig D, Nilson G. 2003. ''True Vipers: Natural History and Toxinology of Old World Vipers''. Krieger Publishing Comp ...

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

to northern and central

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...

and adjacent

Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...

Slovenia Slovenia ( ; sl, Slovenija ), officially the Republic of Slovenia (Slovene: , abbr.: ''RS''), is a country in Central Europe. It is bordered by Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the southeast, an ...

, and

Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...

.Mallow D, Ludwig D, Nilson G. 2003. ''True Vipers: Natural History and Toxinology of Old World Vipers''. Malabar, Florida: Krieger Publishing Company, Malabar. 359 pp. .

Etymology

The

subspecific name In zoological nomenclature, a subspecific name is the third part of a trinomen. In zoology there is only one rank below that of species, namely "subspecies". In botanical nomenclature, there are several levels of subspecific names, such as ''vari ...

, ''francisciredi'', is in honor of

Francesco Redi Francesco Redi (18 February 1626 – 1 March 1697) was an Italian physician, naturalist, biologist, and poet. He is referred to as the "founder of experimental biology", and as the "father of modern parasitology". He was the first person to cha ...

, an

Italian Italian(s) may refer to: * Anything of, from, or related to the people of Italy over the centuries ** Italians, an ethnic group or simply a citizen of the Italian Republic or Italian Kingdom ** Italian language, a Romance language *** Regional Ita ...

physician A physician (American English), medical practitioner (Commonwealth English), medical doctor, or simply doctor, is a health professional who practices medicine, which is concerned with promoting, maintaining or restoring health through th ...

who was a pioneer in experimental

toxinology A toxin is a naturally occurring organic poison produced by metabolic activities of living cells or organisms. Toxins occur especially as a protein or conjugated protein. The term toxin was first used by organic chemist Ludwig Brieger (1849–19 ...

Description

The head is distinctly swollen behind the eyes and upper lips; in adults, this feature is clearly visible when viewed from above. Regarding the color pattern, many examples of this subspecies have a white or cream-colored spot near the outer edge of each

ventral scale In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...

; these spots are smaller in '' V. a. aspis'' (if at all present), but much more apparent in ''V. a. francisciredi''.

Geographic range

Street (1979) describes this subspecies as inhabiting most of

, where it is the most common and widely distributed venomous snake. It also occurs in the

Swiss Swiss may refer to: * the adjectival form of Switzerland * Swiss people Places * Swiss, Missouri * Swiss, North Carolina *Swiss, West Virginia * Swiss, Wisconsin Other uses *Swiss-system tournament, in various games and sports *Swiss Internation ...

canton of

Ticino Ticino (), sometimes Tessin (), officially the Republic and Canton of Ticino or less formally the Canton of Ticino,, informally ''Canton Ticino'' ; lmo, Canton Tesin ; german: Kanton Tessin ; french: Canton du Tessin ; rm, Chantun dal Tessin . ...

, south of the

Monte Ceneri Pass Monte Ceneri is a mountain pass in the canton of Ticino in Switzerland. It connects the Magadino Plain and the Vedeggio Valley across the Lugano Prealps at an elevation of above sea level. It provides the most direct route between the cities ...

, but is not found west of nearby

Lake Maggiore Lake Maggiore (, ; it, Lago Maggiore ; lmo, label=Western Lombard, Lagh Maggior; pms, Lagh Magior; literally 'Greater Lake') or Verbano (; la, Lacus Verbanus) is a large lake located on the south side of the Alps. It is the second largest la ...

. Also in the north, in

Trentino-Alto Adige/Südtirol it, Trentino (man) it, Trentina (woman) or it, Altoatesino (man) it, Altoatesina (woman) or it, Sudtirolesegerman: Südtiroler (man)german: Südtirolerin (woman) , population_note = , population_blank1_title = Official ...

, it can be found as far north as

Meran Merano (, , ) or Meran () is a city and ''comune'' in South Tyrol, northern Italy. Generally best known for its spa resorts, it is located within a basin, surrounded by mountains standing up to above sea level, at the entrance to the Passeier ...

, but is not found in

Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...

, and as far east as

Gorizia Gorizia (; sl, Gorica , colloquially 'old Gorizia' to distinguish it from Nova Gorica; fur, label= Standard Friulian, Gurize, fur, label= Southeastern Friulian, Guriza; vec, label= Bisiacco, Gorisia; german: Görz ; obsolete English ''Gorit ...

(on the border with

). In the south of Italy, it is absent from

Basilicata it, Lucano (man) it, Lucana (woman) , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = , demographics1_footnotes = , demographics1_title1 = , demographics1_info1 = ...

and

Calabria , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = , demographics1_footnotes = , demographics1_title1 = , demographics1_info1 = , demographics1_title2 ...

. In the former

Yugoslavia Yugoslavia (; sh-Latn-Cyrl, separator=" / ", Jugoslavija, Југославија ; sl, Jugoslavija ; mk, Југославија ;; rup, Iugoslavia; hu, Jugoszlávia; rue, label=Pannonian Rusyn, Югославия, translit=Juhoslavija ...

, it is a rare inhabitant of the

Julian Alps The Julian Alps ( sl, Julijske Alpe, it, Alpi Giulie, , ) are a mountain range of the Southern Limestone Alps that stretch from northeastern Italy to Slovenia, where they rise to 2,864 m at Mount Triglav, the highest peak in Slovenia. A large pa ...

(Pozzi, 1966). There have been reports of specimens from

Ripanj Ripanj ( sr-Cyrl, Рипањ) is a suburban settlement of Belgrade, Serbia. It is located in Belgrade's municipality of Voždovac. It has a distinction of being "the largest village of Serbia" taking in account its number of population, but also b ...

near

Belgrade Belgrade ( , ;, ; Names of European cities in different languages: B, names in other languages) is the Capital city, capital and List of cities in Serbia, largest city in Serbia. It is located at the confluence of the Sava and Danube rivers a ...

, Jahorin in

Bosnia Bosnia and Herzegovina ( sh, / , ), abbreviated BiH () or B&H, sometimes called Bosnia–Herzegovina and often known informally as Bosnia, is a country at the crossroads of south and southeast Europe, located in the Balkans. Bosnia and He ...

and other parts of "Yugoslavia", but some of these may have been confused with '' V. berus bosniensis''.Street D. 1979. ''The Reptiles of Northern and Central Europe''. London: B.T. Batsford Ltd. 268 pp. . Nevertheless, the European Molecular Biology Laboratory (EMBL) describes the range of this subspecies as stretching from southern

, northern and central

, to

and northwestern

Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...

. The type locality listed by

Laurenti Laurenti is a surname. It may refer to: People *Adolfo Laurenti (1856-1944), Italian sculptor *Camillo Laurenti (1861–1938), Italian cardinal *Cesare Laurenti (engineer), Italian designer of submarines influencing HMS ''Swordfish'' (1916) *Cesar ...

(1768) for ''Vipera Francisci Redi'' (= ''V. a. francisciredi'') is ''"Austriaco & Italico"'' (=

and

Conservation status

The whole species is classified as Least Concern (LC) according to the

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

Red List of Threatened Species (v3.1, 2008).''Vipera aspis''
a
IUCN Red List
Accessed 31 July 2020. This subspecies is however considered Endangered (EN) in Switzerland''OFEFP red list''
Accessed 31 July 2020. due to its very limited occupation area.

References

External links