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Victoria A. Braithwaite (19 July 1967 – 30 September 2019) was a British scientist who was a Professor of Animal Behaviour and Cognition at
Pennsylvania State University The Pennsylvania State University (Penn State or PSU) is a Public university, public Commonwealth System of Higher Education, state-related Land-grant university, land-grant research university with campuses and facilities throughout Pennsylvan ...
. She was the first person to demonstrate that fish feel pain, which impacted
animal welfare Animal welfare is the well-being of non-human animals. Formal standards of animal welfare vary between contexts, but are debated mostly by animal welfare groups, legislators, and academics. Animal welfare science uses measures such as longevity ...
research and changed guidelines for the treatment of fish in laboratories and fisheries in the UK, Europe, and Canada.


Early life and education

Braithwaite was born in
Bradford Bradford is a city and the administrative centre of the City of Bradford district in West Yorkshire, England. The city is in the Pennines' eastern foothills on the banks of the Bradford Beck. Bradford had a population of 349,561 at the 2011 ...
. She was one of seven children. Her mother, June Pickles, was chair of the Halifax magistrates' bench, and her father, Alan Braithwaite, was a company director. She attended
Bradford Grammar School Bradford Grammar School (BGS) is a co-educational independent day school located in Frizinghall, Bradford, West Yorkshire, England. Entrance is by examination, except for the sixth form, where admission is based on GCSE results. The school ...
, where she was not encouraged to apply to the
University of Oxford , mottoeng = The Lord is my light , established = , endowment = £6.1 billion (including colleges) (2019) , budget = £2.145 billion (2019–20) , chancellor ...
. She did not take this advice, and eventually completed her undergraduate degree in zoology and doctoral degree, supported by a Christopher Welch scholarship, at the
Somerville College, Oxford Somerville College, a constituent college of the University of Oxford in England, was founded in 1879 as Somerville Hall, one of its first two women's colleges. Among its alumnae have been Margaret Thatcher, Indira Gandhi, Dorothy Hodgkin, Ir ...
. Her thesis considered the visual information used by
homing pigeons The homing pigeon, also called the mail pigeon or messenger pigeon, is a variety of domestic pigeons (''Columba livia domestica'') derived from the wild rock dove, selectively bred for its ability to find its way home over extremely long dist ...
to return to their nests. She demonstrated that birds fly home faster when they see their surroundings before being released. She joined the
University of Glasgow , image = UofG Coat of Arms.png , image_size = 150px , caption = Coat of arms Flag , latin_name = Universitas Glasguensis , motto = la, Via, Veritas, Vita , ...
where she studied the cognitive abilities of
salmon Salmon () is the common name for several list of commercially important fish species, commercially important species of euryhaline ray-finned fish from the family (biology), family Salmonidae, which are native to tributary, tributaries of the ...
. It was during this project that she became interested in animal behaviour.


Research and career

Braithwaite spent her career investigating the cognitive abilities of fish and other animals. She showed that fish which live in stable environments (like ponds) use visual markers for guides, whereas fish that live in rivers learn directions using sequences (like left, right, left, right). Her research was the first to show that an animal's cognition is shaped by the worlds in which they live. To recognise her contributions to our understanding of animal navigation, Braithwaite was elected to the Royal Institute of Navigation in 2005. She moved to the
University of Edinburgh The University of Edinburgh ( sco, University o Edinburgh, gd, Oilthigh Dhùn Èideann; abbreviated as ''Edin.'' in post-nominals) is a public research university based in Edinburgh, Scotland. Granted a royal charter by King James VI in 15 ...
in 1995 where she began to investigate whether or not fish felt pain – so-called
nociception Nociception (also nocioception, from Latin ''nocere'' 'to harm or hurt') is the sensory nervous system's process of encoding noxious stimuli. It deals with a series of events and processes required for an organism to receive a painful stimulus, co ...
. Whilst it was well understood that chickens and sheep experienced pain, before Braithwaite it was not known whether or not fish did too. She demonstrated fish felt pain using a series of experiments, the first of which included showing that fish contained the correct anatomy to detect pain (
nociceptors A nociceptor ("pain receptor" from Latin ''nocere'' 'to harm or hurt') is a sensory neuron that responds to damaging or potentially damaging stimuli by sending "possible threat" signals to the spinal cord and the brain. The brain creates the sens ...
). She showed that fish produce pain-killing
opioids Opioids are substances that act on opioid receptors to produce morphine-like effects. Medically they are primarily used for pain relief, including anesthesia. Other medical uses include suppression of diarrhea, replacement therapy for opioid use ...
in the same way that mammals do. She then investigated whether or not they responded to stimuli, and demonstrated that these receptors feel bodily damage and that fish behaviour is different when they are exposed to an unpleasant stimulus. She then demonstrated that fish that had been exposed to an unpleasant stimulus, for example, vinegar, behaved differently when made to perform tasks after they had taken human painkillers. Her work changed protocols regulating the ways fish are treated in laboratories and fisheries in the UK, Europe, and Canada. As of 2019 other countries were also considering making such changes. Braithwaite joined
Pennsylvania State University The Pennsylvania State University (Penn State or PSU) is a Public university, public Commonwealth System of Higher Education, state-related Land-grant university, land-grant research university with campuses and facilities throughout Pennsylvan ...
in 2007, where she was appointed Professor of Fisheries and Biology. Here she demonstrated that providing a more natural environment for fish impacted their cognitive abilities and likelihood to survive in the world. Before Braithwaite's work, fish that had been reared in tanks did not survive when they were released into the world. Whilst a visiting fellow at the
Berlin Institute for Advanced Study The Institute for Advanced Study in Berlin (german: Wissenschaftskolleg zu Berlin) is an interdisciplinary institute founded in 1981 in Grunewald, Berlin, Germany, dedicated to research projects in the natural and social sciences. It is modeled ...
in 2015, Braithwaite analysed the inferences that could be made about animal pain and other emotional states. She was offered the directorship of the IGB Leibniz-Institute of Freshwater Ecology and Inland Fisheries in 2018, but was unable to assume the position.


Selected publications

Her publications include: * * * *


Awards and honours

* 2005 Fellow of the Royal Institute of Navigation * 2006
Fisheries Society of the British Isles The Fisheries Society of the British Isles is an international, non-political, learned society, based in the United Kingdom, that supports scientific activity in fish biology and management through charitable sponsorship. Membership is open to anyo ...
Medal A medal or medallion is a small portable artistic object, a thin disc, normally of metal, carrying a design, usually on both sides. They typically have a commemorative purpose of some kind, and many are presented as awards. They may be int ...
* 2016 Fellow of the
Linnean Society of London The Linnean Society of London is a learned society dedicated to the study and dissemination of information concerning natural history, evolution, and taxonomy. It possesses several important biological specimen, manuscript and literature colle ...


Personal life

Braithwaite in 1992 married and eventually divorced the evolutionary biologist Andrew Read. Together they had two sons. In 2015 she was diagnosed with
pancreatic cancer Pancreatic cancer arises when cell (biology), cells in the pancreas, a glandular organ behind the stomach, begin to multiply out of control and form a Neoplasm, mass. These cancerous cells have the malignant, ability to invade other parts of t ...
. She died on 30 September 2019.


References

{{DEFAULTSORT:Braithwaite, Victoria Women ethologists Alumni of Somerville College, Oxford Alumni of the University of Edinburgh Academics of the University of Glasgow Pennsylvania State University faculty 1967 births 2019 deaths Deaths from pancreatic cancer Fellows of the Linnean Society of London Scientists from Bradford Deaths from cancer in England