Vertigo Moulinsiana
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Desmoulin's whorl snail (''Vertigo moulinsiana'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of minute air-breathing land snail, a
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
pulmonate gastropod mollusc or
micromollusc A micromollusk is a shelled mollusk which is extremely small, even at full adult size. The word is usually, but not exclusively, applied to marine mollusks, although in addition, numerous species of land snails and freshwater mollusks also r ...
in the family
Vertiginidae Vertiginidae, common name the whorl snails, is a family of minute, air-breathing land snails, terrestrial pulmonate gastropod molluscs or micromollusks in the superfamily Pupilloidea. Distribution The distribution of the Vertiginidae is in the ...
, the whorl snails. This species was named in honor of the early-19th-century French naturalist
Charles des Moulins Charles Des Moulins, full name Charles Robert Alexandre Des Moulins (13 March 1798 – 23 December 1875) was a French naturalist, a botanist and malacologist. He was a member of several learned societies, including the American Philosophical ...
.


Habitat

Desmoulin's whorl snail inhabits calcareous wetlands, where there are tall
sedge The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus '' Carex'' ...
s, saw-sedge (''
Cladium mariscus ''Cladium mariscus'' is a species of flowering plant in the sedge family known by the common names swamp sawgrass, great fen-sedge, saw-sedge or sawtooth sedge. Previously it was known as elk sedge. It is native of temperate Europe and Asia wher ...
''), reed-grass (''
Glyceria maxima ''Glyceria maxima'' (syn. ''G. aquatica'' (L.) Wahlenb.; ''G. spectabilis'' Mert. & W.D.J. Koch; ''Molinia maxima'' Hartm.; ''Poa aquatica'' L.), commonly known as great manna grass, reed mannagrass, reed sweet-grass, and greater sweet-grass is a ...
'') or the reed ''
Phragmites australis ''Phragmites australis'', known as the common reed, is a species of plant. It is a broadly distributed wetland grass that can grow up to tall. Description ''Phragmites australis'' commonly forms extensive stands (known as reed beds), which may ...
''.


Distribution

The distribution of this species is Atlantic (the part of the Palearctic area which is under the direct climatic influence of the Atlantic Ocean), and southern-European. This small snail occurs across Europe as far north as southern Sweden. Within
Western Europe Western Europe is the western region of Europe. The region's countries and territories vary depending on context. The concept of "the West" appeared in Europe in juxtaposition to "the East" and originally applied to the ancient Mediterranean ...
, only the populations in England (
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It i ...
) and
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
are considered to be viable, although further populations exist in the
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...
(critically endangered, occupying the White Carpathians Biosphere Reserve, Kokořínsko Landscape Protected Area and Southern Moravia), in
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populou ...
(critically endangered) and elsewhere in Europe (for example:
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan area ...
). Its conservation status in the Czech Republic in 2004–2006 was described as favourable (FV) in the report for the
European Commission The European Commission (EC) is the executive of the European Union (EU). It operates as a cabinet government, with 27 members of the Commission (informally known as "Commissioners") headed by a President. It includes an administrative body ...
in accordance with the Habitats Directive. Its conservation status in
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
is endangered and it occurs in two localities only: near Estañá lake and near Lake of Banyoles.Ramos M. A. (1998). "Implementing the habitats directive for mollusc species in Spain". '' Journal of Conchology'', Molluscan Conservation: A Strategy for the 21st Century, Special Publication 2: 125–132. It is also found in Belgium, Switzerland,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
,
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
,
Slovakia Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the s ...
, Hungary, Romania, Denmark, Norway, Finland, Latvia, Lithuania, Estonia, Belarus,
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...
(
Volhynia Volhynia (also spelled Volynia) ( ; uk, Воли́нь, Volyn' pl, Wołyń, russian: Волы́нь, Volýnʹ, ), is a historic region in Central and Eastern Europe, between south-eastern Poland, south-western Belarus, and western Ukraine. The ...
, critically endangered; around 2014 extinct in
Crimea Crimea, crh, Къырым, Qırım, grc, Κιμμερία / Ταυρική, translit=Kimmería / Taurikḗ ( ) is a peninsula in Ukraine, on the northern coast of the Black Sea, that has been occupied by Russia since 2014. It has a pop ...
),Balashov I. & Gural-Sverlova N. 2012. An annotated checklist of the terrestrial molluscs of Ukraine. ''Journal of Conchology''. 41 (1): 91–109.Balashov I., Yarotskaya M., Filatova J., Starichenko I., Kovalov V. 2017
Terrestrial molluscs of the Tsyr-Pripyat area in Volyn (Northern Ukraine): the first findings of the threatened snail ''Vertigo moulinsiana'' in mainland Ukraine
''Vestnik Zoologii''. 51 (3): 251–258.
Russia,
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
and
Azerbaijan Azerbaijan (, ; az, Azərbaycan ), officially the Republic of Azerbaijan, , also sometimes officially called the Azerbaijan Republic is a transcontinental country located at the boundary of Eastern Europe and Western Asia. It is a part of t ...
. Its distribution also include Algeria and
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria t ...
, but it is possibly extinct in Algeria. This species is mentioned in Annex II of the European Union's
Habitats Directive The Habitats Directive (more formally known as Council Directive 92/43/EEC on the Conservation of natural habitats and of wild fauna and flora) is a directive adopted by the European Community in 1992 as a response to the Berne Convention. The E ...
.


Status in the United Kingdom

In the United Kingdom, Desmoulin's whorl snail is listed as endangered, although it occurs in a number of areas in a band from
Norfolk Norfolk () is a ceremonial and non-metropolitan county in East Anglia in England. It borders Lincolnshire to the north-west, Cambridgeshire to the west and south-west, and Suffolk to the south. Its northern and eastern boundaries are the No ...
to
Dorset Dorset ( ; archaically: Dorsetshire , ) is a county in South West England on the English Channel coast. The ceremonial county comprises the unitary authority areas of Bournemouth, Christchurch and Poole and Dorset. Covering an area of , ...
, with outlying populations in
Kent Kent is a county in South East England and one of the home counties. It borders Greater London to the north-west, Surrey to the west and East Sussex to the south-west, and Essex to the north across the estuary of the River Thames; it faces ...
and the
Llŷn Peninsula The Llŷn Peninsula ( cy, Penrhyn Llŷn or , ) extends into the Irish Sea from North West Wales, south west of the Isle of Anglesey. It is part of the historic county of Caernarfonshire, and historic region and local authority area of Gwynedd. Mu ...
in
North Wales North Wales ( cy, Gogledd Cymru) is a region of Wales, encompassing its northernmost areas. It borders Mid Wales to the south, England to the east, and the Irish Sea to the north and west. The area is highly mountainous and rural, with Snowdonia N ...
and has probably been under-reported in the past because of its minute size. Its presence on the site of the planned
Newbury bypass The Newbury bypass, officially known as The Winchester-Preston Trunk Road (A34) (Newbury Bypass), is a stretch of dual carriageway road which bypasses the town of Newbury in Berkshire, England. It is located to the west of the town and forms p ...
caused the building of that road to be postponed; the building works were able to go ahead once the snails had been moved to a new habitat nearby. It is reported to have since died out at the new site, but the same report states "Desmoulin's whorl snail is now considered less scarce than it was 10 years ago".


Shell description

The
shell Shell may refer to: Architecture and design * Shell (structure), a thin structure ** Concrete shell, a thin shell of concrete, usually with no interior columns or exterior buttresses ** Thin-shell structure Science Biology * Seashell, a hard o ...
is dextral, minute, ovate, ventricose, obtuse at the
apex The apex is the highest point of something. The word may also refer to: Arts and media Fictional entities * Apex (comics), a teenaged super villainess in the Marvel Universe * Ape-X, a super-intelligent ape in the Squadron Supreme universe *Apex, ...
, smoothish, subperforate. The
aperture In optics, an aperture is a hole or an opening through which light travels. More specifically, the aperture and focal length of an optical system determine the cone angle of a bundle of rays that come to a focus in the image plane. An ...
is semiovate, four-toothed, with one tooth on the parietal wall, another on the
columella Lucius Junius Moderatus Columella (; Arabic: , 4 – ) was a prominent writer on agriculture in the Roman Empire. His ' in twelve volumes has been completely preserved and forms an important source on Roman agriculture, together with the wo ...
, and two palatals, the lower one longer. The shell has four
whorls A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). Whorls in nature File:Photograph and axial plane floral ...
, parted by a distinct suture, the last doubly larger than all the others together. It is rather solid, glossy, subpellucid and of a uniform fulvous color. The shell of this species reaches about 3 mm in length. The shell is yellowish or brownish and translucent.


On a stamp

Deutsche Post The Deutsche Post AG, operating under the trade name Deutsche Post DHL Group, is a German multinational package delivery and supply chain management company headquartered in Bonn, Germany. It is one of the world's largest courier companies. ...
featured ''V. moulinsiana'' on a 2002
German German(s) may refer to: * Germany (of or related to) ** Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ge ...
The euro sign () is the currency sign used for the euro, the official currency of the eurozone and unilaterally adopted by Kosovo and Montenegro. The design was presented to the public by the European Commission on 12 December 1996. It consists ...
0.51
postage stamp A postage stamp is a small piece of paper issued by a post office, postal administration, or other authorized vendors to customers who pay postage (the cost involved in moving, insuring, or registering mail), who then affix the stamp to the f ...
as part of a series on endangered species of animals.


References

This article incorporates public domain text from reference. Pilsbry H. A. & Cooke C. M. (1918–1920). ''
Manual of Conchology George Washington Tryon Jr. (20 May 1838 – 5 February 1888) was an American malacologist who worked at the Academy of Natural Sciences in Philadelphia. Biography George Washington Tryon was the son of Edward K. Tryon and Adeline Savidt. ...
. Second series: Pulmonata.'
Volume 25
Pupillidae (Gastrocoptinae, Vertigininae). Philadelphia. pag
178


External links


''Vertigo moulinsiana''
at
Encyclopedia of Life The ''Encyclopedia of Life'' (''EOL'') is a free, online encyclopedia intended to document all of the 1.9 million living species known to science. It is compiled from existing trusted databases curated by experts and with the assistance of ...

''Vertigo moulinsiana''
at
Animalbase AnimalBase is a project brought to life in 2004 and is maintained by the University of Göttingen, Germany. The goal of the AnimalBase project is to digitize early zoological literature, provide copyright-free open access to zoological works, and pr ...
taxonomy,short description, distribution, biology,status (threats), images
Close-up photo of the shell of ''Vertigo moulinsiana''
{{Taxonbar, from=Q811137 Vertigo (gastropod) Gastropods described in 1849