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''Vernonia'' is a genus of about 350
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
forb A forb or phorb is an herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in biology and in vegetation ecology, especially in relation to grasslands and understory. Typically these are dicots without woo ...
s and
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s in the Daisy family
Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae w ...
. Some species are known as ironweed. Some species are edible and of
economic value In economics, economic value is a measure of the benefit provided by a good or service to an economic agent. It is generally measured through units of currency, and the interpretation is therefore "what is the maximum amount of money a specif ...
. They are known for having intense
purple Purple is any of a variety of colors with hue between red and blue. In the RGB color model used in computer and television screens, purples are produced by mixing red and blue light. In the RYB color model historically used by painters, pu ...
flowers. There have been numerous distinct subgenera and subsections named in this
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
, and some botanists have divided the genus into several distinct genera. For instance, the ''
Flora of North America The ''Flora of North America North of Mexico'' (usually referred to as ''FNA'') is a multivolume work describing the native plants and naturalized plants of North America, including the United States, Canada, St. Pierre and Miquelon, and Greenla ...
'' recognizes only about twenty species in ''Vernonia'' ''
sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...
'', seventeen of which are in North America north of Mexico, with the others being found in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
.


Taxonomy

The genus was
circumscribed In geometry, the circumscribed circle or circumcircle of a polygon is a circle that passes through all the vertices of the polygon. The center of this circle is called the circumcenter and its radius is called the circumradius. Not every polyg ...
by
Johann Christian Daniel von Schreber Johann Christian Daniel von Schreber (17 January 1739 in Weißensee, Thuringia – 10 December 1810 in Erlangen), often styled J.C.D. von Schreber, was a German naturalist. Career He was appointed professor of'' materia medica'' at the Univers ...
in Gen. Pl. ed. 8 vol.2 on page 541 in 1791. The genus name of ''Vernonia'' is in honour of William Vernon (1666/67 - ca.1711), who was an English plant collector, ( bryologist) and
entomologist Entomology () is the scientific study of insects, a branch of zoology. In the past the term "insect" was less specific, and historically the definition of entomology would also include the study of animals in other arthropod groups, such as arach ...
from
Cambridge University , mottoeng = Literal: From here, light and sacred draughts. Non literal: From this place, we gain enlightenment and precious knowledge. , established = , other_name = The Chancellor, Masters and Schola ...
, who collected in
Maryland Maryland ( ) is a state in the Mid-Atlantic region of the United States. It shares borders with Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; and Delaware and the Atlantic Ocean to ...
, USA in 1698.


Species

Species of this genus are found in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
,
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
,
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
, and
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. ''Vernonia'' species are well known for hybridizing between similar species in areas of overlapping ranges. There are approximately 350 species in the genus. A selected list is given below.


North America

*''
Vernonia acaulis ''Vernonia acaulis'' is a species of perennial plant from family Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonl ...
'' *''
Vernonia arkansana ''Vernonia arkansana'' (also known as Arkansas ironweed and great ironweed) is a species of perennial plant from family Asteraceae found in south-central United States. The plant is high and wide. The flowers bloom from August to September and ...
'' *''
Vernonia angustifolia ''Vernonia angustifolia'' is a species of flowering plant in the family Asteraceae, native to the southeastern United States (Alabama, Florida, Georgia, Louisiana, Mississippi, North Carolina and South Carolina). It was first described by André ...
'' *''
Vernonia baldwinii ''Vernonia baldwinii'', commonly known as western ironweed or Baldwin's ironweed, is a perennial herb native to the central United States. It is in the Asteraceae (aster) family. Description ''Vernonia baldwinii'' is a tall, perennial herb with ...
'' *''
Vernonia blodgettii ''Vernonia blodgettii'', the Florida ironweed or Blodgett's ironweed, is a species of perennial plant from family Asteraceae that is native to Florida and the Bahamas The Bahamas (), officially the Commonwealth of The Bahamas, is an isla ...
'' *''
Vernonia fasciculata ''Vernonia fasciculata'', the smooth ironweed or common ironweed, or prairie ironweed is a species of perennial plant from family Asteraceae. It is native to Manitoba in Canada and the north-central U.S.A. ''Vernonia fasciculata'' inhabits areas ...
'' *''
Vernonia flaccidifolia ''Vernonia flaccidifolia'' is a species of flowering plant in the family Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. ...
'' *''
Vernonia gigantea ''Vernonia gigantea'' (also known as giant ironweed, tall ironweed or ironweed) is a species of perennial plant from family Asteraceae found in United States and Canada. The plant is native to the eastern United States, north to New York state an ...
'' or ''Vernonia altissima'' *''
Vernonia glauca ''Vernonia glauca'' is a species of flowering plant in the family Asteraceae. It is endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zo ...
'' *''
Vernonia larseniae ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia lettermannii ''Vernonia lettermannii'', the narrowleaf ironweed, is a plant species known only from Arkansas and Oklahoma. It grows on floodplains and terraces at elevations of . ''Vernonia lettermannii'' is a perennial herb up to tall. Leaves are filiform ...
'' *''
Vernonia lindheimeri ''Vernonia lindheimeri'', commonly known as woolly ironweed, is a species of flowering plant in the aster family (Asteraceae). It is native North America, where it is native to the state of Coahuila in Mexico and to the state of Texas in the Uni ...
'' *''
Vernonia marginata ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *'' Vernonia missurica'' *''
Vernonia noveboracensis ''Vernonia noveboracensis'', the New York ironweed or vein-leaf hawkweed, is a plant in the family Asteraceae. It is native to the eastern United States, from Florida to Massachusetts and west to Tennessee, Alabama, and West Virginia and to sout ...
'' *''
Vernonia proctorii ''Vernonia proctorii'' is a rare species of flowering plant in the aster family known by the common name Proctor's ironweed. It is endemic to Puerto Rico, where there are two populations. It is threatened by the destruction of its habitat. It is ...
'' *''
Vernonia pulchella ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia texana ''Vernonia texana'', commonly called Texas ironweed, is a species of flowering plant in the aster family (Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 ge ...
''


South America

*''
Vernonia nonoensis ''Vernonia nonoensis'' is a species of flowering plant in the family Asteraceae that is endemic to Ecuador. Its natural habitat is subtropical or tropical moist montane forest Montane ecosystems are found on the slopes of mountains. The alpi ...
'' *''
Vernonia patens ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia scorpioides ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia condensata ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
''


Africa

*''
Vernonia amygdalina ''Vernonia amygdalina'', a member of the daisy family, is a small shrub that grows in tropical Africa. ''V. amygdalina'' typically grows to a height of . The leaves are elliptical and up to long. Its bark is rough. ''V. amygdalina'' is common ...
'' *'' Vernonia calvoana'' *''
Vernonia colorata ''Vernonia colorata'' is a species of plant in the family Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly refer ...
'' *''
Vernonia galamensis ''Vernonia galamensis'' is a plant in the sunflower family, known for its use as an oilseed. This species, often called ironweed, is the largest source of vernonia oil, which is rich in a useful epoxy fatty acid called vernolic acid and is us ...
'' *''
Vernonia kotschyana ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia staehelinoides ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia cineria ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
vernonia myriantha ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
''


Asia

*''
Vernonia arborea ''Vernonia arborea'' is a species of mid-level rainforest tree in the aster family (Asteraceae, or Compositae) that is found in Western Ghats of India and Sri Lanka and is also widespread in the East Indies. It is the tallest of all composite t ...
'' *''
Vernonia cockburniana ''Vernonia cockburniana'' is a species of flowering plant in the Asteraceae family. It is endemic to Yemen. Its natural habitats are subtropical or tropical dry forests and subtropical or tropical dry shrubland Shrubland, scrubland, scrub, br ...
'' *''
Vernonia elaeagnifolia ''Tarlmounia'' is a plant genus in the family Asteraceae. The sole species is ''Tarlmounia elliptica'' (syn. ''Vernonia elliptica'' and ''V. elaeagnifolia''), native to India, Burma and Thailand and naturalised in southern Taiwan and Queensland ...
'' *''
Vernonia unicata ''Vernonia unicata'' is a species of flowering plant in the aster family that is endemic to Yemen. Its natural habitat is subtropical or tropical dry forest A forest is an area of land dominated by trees. Hundreds of definitions of fores ...
'' *''
Vernonia zollingerianoides ''Vernonia zollingerianoides'' is a species of flowering plant in the family Asteraceae. It is endemic to Java in Indonesia and considered a vulnerable species on the IUCN Red List The International Union for Conservation of Nature (IUCN) Re ...
''


Uses

Several species of ''Vernonia'', including '' V. calvoana'', '' V. amygdalina'', and '' V. colorata'', are eaten as
leaf vegetable Leaf vegetables, also called leafy greens, pot herbs, vegetable greens, or simply greens, are plant leaves eaten as a vegetable, sometimes accompanied by tender petioles and shoots. Leaf vegetables eaten raw in a salad can be called salad gre ...
s. Common names for these species include bitterleaf, onugbu in the
Igbo language Igbo ( , ; Igbo: ''Ásụ̀sụ́ Ìgbò'' ) is the principal native language cluster of the Igbo people, a meta-ethnicity from Southeastern Nigeria. The number of Igboid languages depends on how one classifies a language versus a dialect, so ...
, ewuro and ndole. They are common in most
West Africa West Africa or Western Africa is the westernmost region of Africa. The United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Mali, Maurit ...
n and
Central Africa Central Africa is a subregion of the African continent comprising various countries according to different definitions. Angola, Burundi, the Central African Republic, Chad, the Democratic Republic of the Congo, the Republic of the Congo, ...
n countries. They are one of the most widely consumed leaf vegetables of
Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...
, where the onugbu soup is a local delicacy of the
Igbo people The Igbo people ( , ; also spelled Ibo" and formerly also ''Iboe'', ''Ebo'', ''Eboe'', * * * ''Eboans'', ''Heebo''; natively ) are an ethnic group in Nigeria. They are primarily found in Abia, Anambra, Ebonyi, Enugu, and Imo States. A ...
, and of
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
, where they are a key ingredient of
Ndolé Ndolé is a Cameroonian dish consisting of stewed nuts, ndoleh (bitter leaves indigenous to West Africa), and fish or beef. The dish may also contain shrimp. It is traditionally eaten with plantains, bobolo (a Cameroonian dish made of fermented ...
: the national dish of
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
. The leaves have a sweet and bitter taste. They are sold fresh or
dried Drying is a mass transfer process consisting of the removal of water or another solvent by evaporation from a solid, semi-solid or liquid. This process is often used as a final production step before selling or packaging products. To be consider ...
and are a typical ingredient in
egusi soup Egusi (Yoruba: '' ẹ̀gúsí,'' Igbo: ègwusi), also known as, agusi, ohue, Ikpan, Ikon, or agushi) is the name for the protein-rich seeds of certain cucurbitaceous plants (squash, melon, gourd), which, after being dried and ground, are used ...
. ''
Vernonia galamensis ''Vernonia galamensis'' is a plant in the sunflower family, known for its use as an oilseed. This species, often called ironweed, is the largest source of vernonia oil, which is rich in a useful epoxy fatty acid called vernolic acid and is us ...
'' is used as an
oilseed Vegetable oils, or vegetable fats, are oils extracted from seeds or from other parts of fruits. Like animal fats, vegetable fats are ''mixtures'' of triglycerides. Soybean oil, grape seed oil, and cocoa butter are examples of seed oils, or fat ...
in
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
. It is grown in many parts of
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
, especially around the city of
Harar Harar ( amh, ሐረር; Harari: ሀረር; om, Adare Biyyo; so, Herer; ar, هرر) known historically by the indigenous as Gey (Harari: ጌይ ''Gēy'', ) is a walled city in eastern Ethiopia. It is also known in Arabic as the City of Saint ...
, with an average seed yield of 2 to 2.5 t/ha. It is reported that the Ethiopian strains of ''Vernonia'' have the highest oil content, up to 41.9% with up to 80%
vernolic acid Vernolic acid (leukotoxin) is a long chain fatty acid that is monounsaturated and contains an epoxide. It is the R,R-''cis'' epoxide derived from the C12–C13 alkene of linoleic acid. Vernolic acid was first definitively characterized in 1954. I ...
, and is used in paint formulations, coatings plasticizers, and as a reagent for many industrial chemicals. ''Vernonia amygdalina'' is used in traditional herbal medicine. These leaves are exported from several African countries and can be purchased in grocery stores aiming to serve African clients. In Brazil, '' V. condensata'' is commonly known as "figatil" or "necroton" and used in local traditional medicine.


Ecology

''Vernonia'' species are used as food plants by the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e of some
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
species including '' Coleophora vernoniaeella'' (which feeds exclusively on the genus) and ''
Schinia regia ''Schinia regia'' is a moth of the family Noctuidae. It is found in North America, including Colorado, Kansas, Nebraska, Oklahoma, New Mexico and Texas. The wingspan The wingspan (or just span) of a bird or an airplane is the distance f ...
'' (which feeds exclusively on '' V. texana''). Vernonia is a very diverse genus, varying from the tiny V. desertorum of
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
which is only three inches (8 centimeters) tall to
Vernonia arborea ''Vernonia arborea'' is a species of mid-level rainforest tree in the aster family (Asteraceae, or Compositae) that is found in Western Ghats of India and Sri Lanka and is also widespread in the East Indies. It is the tallest of all composite t ...
of the
East Indies The East Indies (or simply the Indies), is a term used in historical narratives of the Age of Discovery. The Indies refers to various lands in the East or the Eastern hemisphere, particularly the islands and mainlands found in and around t ...
which, at 117 feet (36 meters) is the tallest of all composites; a 472 fold difference in height. The leaves can vary from quite small up to four feet (1.2 meters) long by up to 15 inches (38 centimeters) in width in the case of V. conferta of
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
. Karsten and Schenk, Vegetationsbilder, (1911) Volume 8 Plate 40 with caption


References

{{Taxonbar, from=Q1411311 * Asteraceae genera