Vernonia Apiculata
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''Vernonia'' is a genus of about 350 species of forbs and
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been numerous distinct subgenera and subsections named in this genus, and some botanists have divided the genus into several distinct genera. For instance, the ''
Flora of North America The ''Flora of North America North of Mexico'' (usually referred to as ''FNA'') is a multivolume work describing the native plants and naturalized plants of North America, including the United States, Canada, St. Pierre and Miquelon, and Greenla ...
'' recognizes only about twenty species in ''Vernonia'' ''
sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...
'', seventeen of which are in North America north of Mexico, with the others being found in South America.


Taxonomy

The genus was
circumscribed In geometry, the circumscribed circle or circumcircle of a polygon is a circle that passes through all the vertices of the polygon. The center of this circle is called the circumcenter and its radius is called the circumradius. Not every polyg ...
by
Johann Christian Daniel von Schreber Johann Christian Daniel von Schreber (17 January 1739 in Weißensee, Thuringia – 10 December 1810 in Erlangen), often styled J.C.D. von Schreber, was a German naturalist. Career He was appointed professor of'' materia medica'' at the Univers ...
in Gen. Pl. ed. 8 vol.2 on page 541 in 1791. The genus name of ''Vernonia'' is in honour of William Vernon (1666/67 - ca.1711), who was an English plant collector, ( bryologist) and
entomologist Entomology () is the scientific study of insects, a branch of zoology. In the past the term "insect" was less specific, and historically the definition of entomology would also include the study of animals in other arthropod groups, such as arach ...
from Cambridge University, who collected in Maryland, USA in 1698.


Species

Species of this genus are found in South America, Africa, Southeast Asia, and
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. ''Vernonia'' species are well known for hybridizing between similar species in areas of overlapping ranges. There are approximately 350 species in the genus. A selected list is given below.


North America

*'' Vernonia acaulis'' *'' Vernonia arkansana'' *'' Vernonia angustifolia'' *'' Vernonia baldwinii'' *'' Vernonia blodgettii'' *'' Vernonia fasciculata'' *'' Vernonia flaccidifolia'' *'' Vernonia gigantea'' or ''Vernonia altissima'' *'' Vernonia glauca'' *''
Vernonia larseniae ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia lettermannii ''Vernonia lettermannii'', the narrowleaf ironweed, is a plant species known only from Arkansas and Oklahoma. It grows on floodplains and terraces at elevations of . ''Vernonia lettermannii'' is a perennial herb up to tall. Leaves are filiform ...
'' *''
Vernonia lindheimeri ''Vernonia lindheimeri'', commonly known as woolly ironweed, is a species of flowering plant in the aster family (Asteraceae). It is native North America, where it is native to the state of Coahuila in Mexico and to the state of Texas in the Uni ...
'' *''
Vernonia marginata ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia missurica ''Vernonia missurica'', the Missouri ironweed, is a species of magenta-flowered perennial plant from family Asteraceae native to the central and east central United States. Description The plant is in height and in width, and in some cases c ...
'' *''
Vernonia noveboracensis ''Vernonia noveboracensis'', the New York ironweed or vein-leaf hawkweed, is a plant in the family Asteraceae. It is native to the eastern United States, from Florida to Massachusetts and west to Tennessee, Alabama, and West Virginia and to sout ...
'' *''
Vernonia proctorii ''Vernonia proctorii'' is a rare species of flowering plant in the aster family known by the common name Proctor's ironweed. It is endemic to Puerto Rico, where there are two populations. It is threatened by the destruction of its habitat. It is ...
'' *''
Vernonia pulchella ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia texana ''Vernonia texana'', commonly called Texas ironweed, is a species of flowering plant in the aster family (Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 ge ...
''


South America

*''
Vernonia nonoensis ''Vernonia nonoensis'' is a species of flowering plant in the family Asteraceae that is endemic to Ecuador. Its natural habitat is subtropical or tropical moist montane forest Montane ecosystems are found on the slopes of mountains. The alpi ...
'' *''
Vernonia patens ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia scorpioides ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia condensata ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
''


Africa

*'' Vernonia amygdalina'' *''
Vernonia calvoana ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have bee ...
'' *''
Vernonia colorata ''Vernonia colorata'' is a species of plant in the family Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly refer ...
'' *''
Vernonia galamensis ''Vernonia galamensis'' is a plant in the sunflower family, known for its use as an oilseed. This species, often called ironweed, is the largest source of vernonia oil, which is rich in a useful epoxy fatty acid called vernolic acid and is us ...
'' *''
Vernonia kotschyana ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia staehelinoides ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
Vernonia cineria ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' *''
vernonia myriantha ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
''


Asia

*'' Vernonia arborea'' *''
Vernonia cockburniana ''Vernonia cockburniana'' is a species of flowering plant in the Asteraceae family. It is endemic to Yemen. Its natural habitats are subtropical or tropical dry forests and subtropical or tropical dry shrubland Shrubland, scrubland, scrub, br ...
'' *''
Vernonia elaeagnifolia ''Tarlmounia'' is a plant genus in the family Asteraceae. The sole species is ''Tarlmounia elliptica'' (syn. ''Vernonia elliptica'' and ''V. elaeagnifolia''), native to India, Burma and Thailand and naturalised in southern Taiwan and Queensland ...
'' *''
Vernonia unicata ''Vernonia unicata'' is a species of flowering plant in the aster family that is endemic to Yemen. Its natural habitat is subtropical or tropical dry forest A forest is an area of land dominated by trees. Hundreds of definitions of fores ...
'' *''
Vernonia zollingerianoides ''Vernonia zollingerianoides'' is a species of flowering plant in the family Asteraceae. It is endemic to Java in Indonesia and considered a vulnerable species on the IUCN Red List The International Union for Conservation of Nature (IUCN) Re ...
''


Uses

Several species of ''Vernonia'', including '' V. calvoana'', '' V. amygdalina'', and '' V. colorata'', are eaten as leaf vegetables. Common names for these species include bitterleaf, onugbu in the Igbo language, ewuro and ndole. They are common in most West African and Central African countries. They are one of the most widely consumed leaf vegetables of Nigeria, where the onugbu soup is a local delicacy of the
Igbo people The Igbo people ( , ; also spelled Ibo" and formerly also ''Iboe'', ''Ebo'', ''Eboe'', * * * ''Eboans'', ''Heebo''; natively ) are an ethnic group in Nigeria. They are primarily found in Abia, Anambra, Ebonyi, Enugu, and Imo States. A ...
, and of Cameroon, where they are a key ingredient of Ndolé: the national dish of Cameroon. The leaves have a sweet and bitter taste. They are sold fresh or
dried Drying is a mass transfer process consisting of the removal of water or another solvent by evaporation from a solid, semi-solid or liquid. This process is often used as a final production step before selling or packaging products. To be consider ...
and are a typical ingredient in egusi soup. ''
Vernonia galamensis ''Vernonia galamensis'' is a plant in the sunflower family, known for its use as an oilseed. This species, often called ironweed, is the largest source of vernonia oil, which is rich in a useful epoxy fatty acid called vernolic acid and is us ...
'' is used as an oilseed in
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
. It is grown in many parts of Ethiopia, especially around the city of Harar, with an average seed yield of 2 to 2.5 t/ha. It is reported that the Ethiopian strains of ''Vernonia'' have the highest oil content, up to 41.9% with up to 80%
vernolic acid Vernolic acid (leukotoxin) is a long chain fatty acid that is monounsaturated and contains an epoxide. It is the R,R-''cis'' epoxide derived from the C12–C13 alkene of linoleic acid. Vernolic acid was first definitively characterized in 1954. I ...
, and is used in paint formulations, coatings plasticizers, and as a reagent for many industrial chemicals. ''Vernonia amygdalina'' is used in traditional herbal medicine. These leaves are exported from several African countries and can be purchased in grocery stores aiming to serve African clients. In Brazil, '' V. condensata'' is commonly known as "figatil" or "necroton" and used in local traditional medicine.


Ecology

''Vernonia'' species are used as food plants by the larvae of some
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
species including ''
Coleophora vernoniaeella ''Coleophora vernoniaeella'' is a moth of the family Coleophoridae. It is found in the United States, including Kentucky and Missouri. The larvae feed on the leaves of ''Vernonia'' and ''Helianthus ''Helianthus'' () is a genus comprising abo ...
'' (which feeds exclusively on the genus) and ''
Schinia regia ''Schinia regia'' is a moth of the family Noctuidae. It is found in North America, including Colorado, Kansas, Nebraska, Oklahoma, New Mexico and Texas. The wingspan The wingspan (or just span) of a bird or an airplane is the distance f ...
'' (which feeds exclusively on '' V. texana''). Vernonia is a very diverse genus, varying from the tiny V. desertorum of Brazil which is only three inches (8 centimeters) tall to Vernonia arborea of the East Indies which, at 117 feet (36 meters) is the tallest of all composites; a 472 fold difference in height. The leaves can vary from quite small up to four feet (1.2 meters) long by up to 15 inches (38 centimeters) in width in the case of V. conferta of Cameroon. Karsten and Schenk, Vegetationsbilder, (1911) Volume 8 Plate 40 with caption


References

{{Taxonbar, from=Q1411311 * Asteraceae genera