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The verdin (''Auriparus flaviceps'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
penduline tit The penduline tits constitute the family, Remizidae, of small passerine birds, related to the true tits. All but the verdin make elaborate bag nests hanging from trees (whence "penduline", hanging), usually over water. Characteristics Penduline ...
. It is the only species in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Auriparus'' and the only representative of the old world family Remizidae to be found in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
.


Taxonomy

The verdin was formally described in 1850 by the Swedish zoologist Carl Jakob Sundevall under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Aegithalus flaviceps''. It is now the only species placed in the genus ''Auriparus'' that was introduced in 1864 by the American naturalist
Spencer Fullerton Baird Spencer Fullerton Baird (; February 3, 1823 – August 19, 1887) was an American naturalist, ornithologist, ichthyologist, Herpetology, herpetologist, and museum curator. Baird was the first curator to be named at the Smithsonian Institution. He ...
. The genus name combines the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''aurum'', ''auri'' meaning "gold" with the genus name '' Parus'' that was introduced by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in 1758 for the tits. The specific epithet ''flaviceps'' combines the Latin ''flavis'' meaning "yellow" with ''-ceps'' meaning "-capped". Six
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised: * ''A. f. acaciarum'' Grinnell, 1931 – southwest USA to north Baja California, central
Sonora Sonora (), officially Estado Libre y Soberano de Sonora ( en, Free and Sovereign State of Sonora), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. The state is d ...
, central
Chihuahua Chihuahua may refer to: Places *Chihuahua (state), a Mexican state **Chihuahua (dog), a breed of dog named after the state **Chihuahua cheese, a type of cheese originating in the state **Chihuahua City, the capital city of the state **Chihuahua Mun ...
and central
Durango Durango (), officially named Estado Libre y Soberano de Durango ( en, Free and Sovereign State of Durango; Tepehuán: ''Korian''; Nahuatl: ''Tepēhuahcān''), is one of the 31 states which make up the 32 Federal Entities of Mexico, situated in ...
(northwest Mexico) * ''A. f. ornatus'' (
Lawrence Lawrence may refer to: Education Colleges and universities * Lawrence Technological University, a university in Southfield, Michigan, United States * Lawrence University, a liberal arts university in Appleton, Wisconsin, United States Preparator ...
, 1851) – south-central USA and northeast Mexico * ''A. f. flaviceps'' (
Sundevall Carl Jakob Sundevall (22 October 1801, Högestad – 2 February 1875) was a Swedish zoologist. Sundevall studied at Lund University, where he became a Ph.D. in 1823. After traveling to East Asia, he studied medicine, graduating as Doctor of Med ...
, 1850) – central Baja California, south Sonora and north
Sinaloa Sinaloa (), officially the Estado Libre y Soberano de Sinaloa ( en, Free and Sovereign State of Sinaloa), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. It is d ...
(northwest Mexico) * ''A. f. lamprocephalus'' Oberholser, 1897 – south Baja California (northwest Mexico) * ''A. f. sinaloae'' Phillips, AR, 1986 – northwest Sinaloa (northwest Mexico) * ''A. f. hidalgensis'' Phillips, AR, 1986 – north-central Mexico


Description

The verdin is a very small bird. At in length, it rivals the
American bushtit The American bushtit (''Psaltriparus minimus'') is the only species placed in the genus ''Psaltriparus'' and the only species in the family Aegithalidae that is found in the New World. In North America, it is referred to simply as "bushtit". Tax ...
as one of the smallest
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
s in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. It is gray overall, and adults have a bright yellow head and rufous shoulder patch (the lesser coverts). Unlike the tits, it has a sharply pointed
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
. Juveniles have a light gray body without the rufous shoulder patch and yellow head. The base of the short bill is thick by the head, and draws to a sharp point at the tip.


Vocalizations

Little is known about geographic or individual variation in vocalizations. Verdins produce a variety of calls, loud for their size. The typical call note is a repetitive "tschep" sound repeated in rapid staccato. In Texas, rapid "chip-chip-chip" calls have been observed, all at the same pitch and at the same speed in each call series.” The day of hatching, nestlings can produce short peep notes, faint and high-pitched. Fledglings, during their post-juvenile molt, can make soft warbling notes, similar to more complex signatures of adult calls. The verdin gives rapid "tschep" calls frequently while foraging. Unmated males sound "tseet" calls repeatedly in early spring, which may function to attract females. "Tseet" calls are also given by either sex of a mated pair, by solitary foraging birds, and by older nestlings. This may be used as an identifier call to help keep members of the breeding pair together as they move through brush to forage.Dawson, W. L. (1923) The Birds of California: A Complete, Scientific and Popular Account of the 580 Species and Subspecies of Birds Found in the State. Volume 2. South Moulton, San Diego, CA, USA "Tweedle" warbling calls may reinforce a breeding pair bond. The verdin also gives a "gee-gee-gee-gee" alarm call in response to predators or when captured.


Distribution and habitat

Two subspecies nest in the United States. The subspecies ''A. f. acaciarum'' is resident from southern California, Nevada, and Utah southward into Mexico; it is associated mostly with the
Sonoran Desert The Sonoran Desert ( es, Desierto de Sonora) is a desert in North America and ecoregion that covers the northwestern Mexican states of Sonora, Baja California, and Baja California Sur, as well as part of the southwestern United States (in Arizona ...
. The subspecies ''A. f. ornatus'' is found from New Mexico and Oklahoma southward into Mexico and is associated with the
Chihuahuan Desert The Chihuahuan Desert ( es, Desierto de Chihuahua, ) is a desert ecoregion designation covering parts of northern Mexico and the southwestern United States. It occupies much of far West Texas, the middle to lower Rio Grande Valley and the lower P ...
and its scrublands. The remaining 4-5 subspecies are resident to Mexico, and aren't known to migrate into the United States. Verdins nest in shrubs and thorny thickets with few trees. Their nests are best described as spherical masses of bulky twigs and branches that are then lined with grasses, feathers, and hair. The entrance to the nest is about 1" in diameter, and is usually at the bottom of the sphere. They feed in shrub-land and scrubby outcrops where insects are hosted and berries and seeds are plentiful.


Behavior and ecology


Breeding

Verdins nest in the spring and tend to form monogamous pairs, but do not stay together year-round. Otherwise, they are usually solitary. Both males and females build nests for roosting and nesting, and both incubate the eggs and tend to young. Clutch sizes vary from 3-6 eggs, which are light green with irregular dark red-brown spots, concentrated at the bottom end of the egg. No formal courtship display is known for the species, but males sing more frequently during breeding and sometimes also build a "display nest" of sorts. This display nest may help to display partner suitability, but this correlation has not been confirmed. Once paired, the male and female claim a territory (as large as 20
acre The acre is a unit of land area used in the imperial Imperial is that which relates to an empire, emperor, or imperialism. Imperial or The Imperial may also refer to: Places United States * Imperial, California * Imperial, Missouri * Imp ...
s) and will defend the boundary starting early in the breeding season through
fledging Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
. After fledging, the young forage with both parents until winter, and then disperse short distances from their hatching site.


Food and feeding

Verdins are
insectivorous A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores were ...
, continuously foraging among the
desert A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...
trees and scrubs. Verdins occasionally try to obtain sugar water from hummingbird feeders. Feeds on many kinds of tiny insects, both adults and larvae, and also consumes hard and soft
mast Mast, MAST or MASt may refer to: Engineering * Mast (sailing), a vertical spar on a sailing ship * Flagmast, a pole for flying a flag * Guyed mast, a structure supported by guy-wires * Mooring mast, a structure for docking an airship * Radio mast ...
of many trees and shrubs.


Conservation status

Although the verdin is listed as a species of least concern Cby the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
, The North American Breeding Bird Survey suggests that verdin populations have declined by 1.9% per year between 1968 and 2015. This decline could mean that the species has experienced a cumulative decrease of 60% in that 47-year period. Threats to verdin populations include habitat loss as a result of
urbanization Urbanization (or urbanisation) refers to the population shift from rural to urban areas, the corresponding decrease in the proportion of people living in rural areas, and the ways in which societies adapt to this change. It is predominantly t ...
, human structure collisions, and
predation Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
.


References


External links


Verdin profile
at State of Utah, Natural Resources, Division of Wildlife Resources * * * * {{Taxonbar, from=Q1300274 Auriparus Native birds of the Southwestern United States Birds of Mexico Endemic birds of Southwestern North America Birds described in 1850