Vegetation is an assemblage of

plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...

species and the

ground cover Groundcover or ground cover is any plant that grows over an area of ground. Groundcover provides protection of the topsoil from erosion and drought. In an ecosystem, the ground cover forms the layer of vegetation below the shrub layer known as t ...

they provide. It is a general term, without specific reference to particular

taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...

, life forms, structure, spatial extent, or any other specific

botanical Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...

or geographic characteristics. It is broader than the term ''

flora Flora is all the plant life present in a particular region or time, generally the naturally occurring (indigenous) native plants. Sometimes bacteria and fungi are also referred to as flora, as in the terms '' gut flora'' or '' skin flora''. E ...

'' which refers to

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

composition. Perhaps the closest

synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...

plant community A plant community is a collection or association of plant species within a designated geographical unit, which forms a relatively uniform patch, distinguishable from neighboring patches of different vegetation types. The components of each plant co ...

, but ''vegetation'' can, and often does, refer to a wider range of spatial scales than that term does, including scales as large as the global. Primeval

redwood forest ''Sequoia sempervirens'' ()''Sunset Western Garden Book,'' 1995:606–607 is the sole living species of the genus '' Sequoia'' in the cypress family Cupressaceae (formerly treated in Taxodiaceae). Common names include coast redwood, coastal ...

s, coastal

mangrove A mangrove is a shrub or tree that grows in coastal saline water, saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evoluti ...

stands,

sphagnum bog A bog or bogland is a wetland that accumulates peat as a deposit of dead plant materials often mosses, typically sphagnum moss. It is one of the four main types of wetlands. Other names for bogs include mire, mosses, quagmire, and muskeg; a ...

desert A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...

soil crusts,

roadside Roadside may refer to: *Road verge, a strip of greenery between a road and a sidewalk *Shoulder (road), an emergency stopping lane by the verge of a road *Roadside, Caithness, Scotland, a village * ''Roadside'' (film), a 2013 American horror film ...

weed patches, wheat fields, cultivated

garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate both ...

s and lawns; all are encompassed by the term ''vegetation''. The

vegetation type Vegetation classification is the process of classifying and mapping the vegetation over an area of the earth's surface. Vegetation classification is often performed by state based agencies as part of land use, resource management, resource and envi ...

is defined by characteristic dominant species, or a common aspect of the assemblage, such as an elevation range or environmental commonality. The contemporary use of ''vegetation'' approximates that of ecologist Frederic Clements' term earth cover, an expression still used by the

Bureau of Land Management The Bureau of Land Management (BLM) is an agency within the United States Department of the Interior responsible for administering federal lands. Headquartered in Washington DC, and with oversight over , it governs one eighth of the country's la ...

History of definition

The distinction between vegetation (the general appearance of a community) and

(the taxonomic composition of a community) was first made by

Jules Thurmann Jules Thurmann (5 November 1804, Neuf-Brisach in Haut-Rhin, France – 25 July 1855, Porrentruy) was an Alsatian French-Swiss geologist and botanist. He studied at the college in Porrentruy, then continued his education at the University of ...

(1849). Prior to this, the two terms (vegetation and flora) were used indiscriminately,Martins, F. R. & Batalha, M. A. (2011). Formas de vida, espectro biológico de Raunkiaer e fisionomia da vegetação. In: Felfili, J. M., Eisenlohr, P. V.; Fiuza de Melo, M. M. R.; Andrade, L. A.; Meira Neto, J. A. A. (Org.). ''Fitossociologia no Brasil: métodos e estudos de caso.'' Vol. 1. Viçosa: Editora UFV. p. 44-85. . Earlier version, 2003, . and still are in some contexts.

Augustin de Candolle Augustin Pyramus (or Pyrame) de Candolle (, , ; 4 February 17789 September 1841) was a Swiss botanist. René Louiche Desfontaines launched de Candolle's botanical career by recommending him at a herbarium. Within a couple of years de Candolle ...

(1820) also made a similar distinction but he used the terms "station" (

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

type) and "habitation" ( botanical region). Later, the concept of vegetation would influence the usage of the term

biome A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...

with the inclusion of the animal element.Coutinho, L. M. (2006). O conceito de bioma. ''Acta Bot. Bras.'' 20(1): 13-23, . Other concepts similar to vegetation are "

physiognomy Physiognomy (from the Greek , , meaning "nature", and , meaning "judge" or "interpreter") is the practice of assessing a person's character or personality from their outer appearance—especially the face. The term can also refer to the general ...

of vegetation" ( Humboldt, 1805, 1807) and "formation" ( Grisebach, 1838, derived from "''Vegetationsform''", Martius, 1824). Departing from

Linnean taxonomy Linnaean taxonomy can mean either of two related concepts: # The particular form of biological classification (taxonomy) set up by Carl Linnaeus, as set forth in his ''Systema Naturae'' (1735) and subsequent works. In the taxonomy of Linnaeus t ...

, Humboldt established a new science, dividing

plant geography Phytogeography (from Greek language, Greek φυτόν, ''phytón'' = "plant" and γεωγραφία, ''geographía'' = "geography" meaning also distribution) or botanical geography is the branch of biogeography that is concerned with the geographi ...

between taxonomists who studied plants as taxa and geographers who studied plants as vegetation. The physiognomic approach in the study of vegetation is common among biogeographers working on vegetation on a world scale, or when there is a lack of taxonomic knowledge of someplace (e.g., in the tropics, where biodiversity is commonly high).Beard J.S. (1978). The Physiognomic Approach. In: R. H. Whittaker (editor). ''Classification of Plant Communities'', pp 33-64

The concept of "

" is more ambiguous. The definition of a specific vegetation type may include not only physiognomy but also floristic and habitat aspects. Furthermore, the phytosociology, phytosociological approach in the study of vegetation relies upon a fundamental unit, the

plant association A plant community is a collection or association of plant species within a designated geographical unit, which forms a relatively uniform patch, distinguishable from neighboring patches of different vegetation types. The components of each plant co ...

, which is defined upon flora. An influential, clear and simple classification scheme for types of vegetation was produced by

Wagner Wilhelm Richard Wagner ( ; ; 22 May 181313 February 1883) was a German composer, theatre director, polemicist, and conductor who is chiefly known for his operas (or, as some of his mature works were later known, "music dramas"). Unlike most op ...

von Sydow von Sydow, often used in German and Swedish with the word "von" meaning "of", is a locational surname, which originally meant simply a person from one of several places called Sydow in northern Germany.''Dictionary of American Family Names''"Von Syd ...

(1888).Wagner, H. & von Sydow, E. 1888. ''Sydow-Wagners methodischer Schulatlas''. Gotha: Perthes, . 23th (last) ed., 1944

. Other important works with a physiognomic approach includes Grisebach (1872), Eugenius Warming, Warming (1895, 1909), Schimper (1898),

Tansley Tansley is a village on the southern edge of the Derbyshire Peak District, two miles east of Matlock. History Tansley is recorded in the Domesday Book as Taneslege, and its name comes from the combination of the Old English elements ''tān'' a ...

and Chipp (1926), Rübel (1930),

Burtt Davy Joseph Burtt Davy (7 March 1870 Findern, Derbyshire – 20 August 1940 Birmingham) was a Quaker botanist and agrostologist. He was the first curator of the Forest Herbarium (FHO) at the Imperial Forestry Institute when it was founded in 1924 u ...

(1938),

Beard A beard is the hair that grows on the jaw, chin, upper lip, lower lip, cheeks, and neck of humans and some non-human animals. In humans, usually pubescent or adult males are able to grow beards. Throughout the course of history, societal at ...

(1944, 1955), André Aubréville (1956, 1957), Trochain (1955, 1957), Küchler (1967), Ellenberg and Mueller-Dombois (1967) (see

vegetation classification Vegetation classification is the process of classifying and mapping the vegetation over an area of the earth's surface. Vegetation classification is often performed by state based agencies as part of land use, resource and environmental management. ...

Classifications

There are many approaches for the classification of vegetation (physiognomy, flora, ecology, etc.).Mueller-Dombois, D. 1984. Classification and Mapping of Plant Communities: a Review with Emphasis on Tropical Vegetation. In: G. M. Woodwell (ed.) ''The Role of Terrestrial Vegetation in the Global Carbon Cycle: Measurement by Remote Sensing'', J. Wiley & Sons, New York, pp. 21-88, . Much of the work on vegetation classification comes from European and North American ecologists, and they have fundamentally different approaches. In North America, vegetation types are based on a combination of the following criteria: climate pattern,

plant habit Habit, equivalent to habitus in some applications in biology, refers variously to aspects of behaviour or structure, as follows: *In zoology (particularly in ethology), habit usually refers to aspects of more or less predictable ''behaviour'', ...

phenology Phenology is the study of periodic events in biological life cycles and how these are influenced by seasonality, seasonal and interannual variations in climate, as well as environmental factor, habitat factors (such as elevation). Examples includ ...

and/or growth form, and dominant species. In the current US standard (adopted by the

Federal Geographic Data Committee The Federal Geographic Data Committee (FGDC) is a United States government committee which promotes the coordinated development, use, sharing, and dissemination of geospatial data on a national basis. Its 32 members are representatives from the E ...

(FGDC), and originally developed by

UNESCO The United Nations Educational, Scientific and Cultural Organization is a specialized agency of the United Nations (UN) aimed at promoting world peace and security through international cooperation in education, arts, sciences and culture. It ...

and

The Nature Conservancy The Nature Conservancy (TNC) is a global environmental organization headquartered in Arlington, Virginia. it works via affiliates or branches in 79 countries and territories, as well as across every state in the US. Founded in 1951, The Natu ...

), the classification is

hierarchical A hierarchy (from Greek: , from , 'president of sacred rites') is an arrangement of items (objects, names, values, categories, etc.) that are represented as being "above", "below", or "at the same level as" one another. Hierarchy is an important ...

and incorporates the non-floristic criteria into the upper (most general) five levels and limited floristic criteria only into the lower (most specific) two levels. In Europe, classification often relies much more heavily, sometimes entirely, on floristic (species) composition alone, without explicit reference to climate, phenology or growth forms. It often emphasizes indicator or diagnostic species which may distinguish one classification from another. In the FGDC standard, the hierarchy levels, from most general to most specific, are: ''system, class, subclass, group, formation, alliance, ''and'' association''. The lowest level, or association, is thus the most precisely defined, and incorporates the names of the dominant one to three (usually two) species of a type. An example of a vegetation type defined at the level of class might be "''Forest, canopy cover > 60%''"; at the level of a formation as "''Winter-rain, broad-leaved, evergreen, sclerophyllous, closed-canopy forest''"; at the level of alliance as "''Arbutus menziesii'' forest"; and at the level of association as "''Arbutus menziesii-Lithocarpus dense flora'' forest", referring to Pacific madrone-tanoak forests which occur in California and Oregon, USA. In practice, the levels of the alliance and/or an association are the most often used, particularly in vegetation mapping, just as the Latin binomial is most often used in discussing particular species in taxonomy and in general communication.

Dynamics

Like all the biological systems, plant communities are temporally and spatially dynamic; they change at all possible scales. Dynamism in vegetation is defined primarily as changes in species composition and/or vegetation structure.

Temporal dynamics

Temporally, a large number of processes or events can cause change, but for sake of simplicity, they can be categorized roughly as either abrupt or gradual. Abrupt changes are generally referred to as disturbances; these include things like

wildfire A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of Combustibility and flammability, combustible vegetation. Depending on the type of vegetation present, a wildfire ...

s, high

winds Wind is the natural movement of air or other gases relative to a planet's surface. Winds occur on a range of scales, from thunderstorm flows lasting tens of minutes, to local breezes generated by heating of land surfaces and lasting a few hou ...

landslide Landslides, also known as landslips, are several forms of mass wasting that may include a wide range of ground movements, such as rockfalls, deep-seated grade (slope), slope failures, mudflows, and debris flows. Landslides occur in a variety of ...

flood A flood is an overflow of water ( or rarely other fluids) that submerges land that is usually dry. In the sense of "flowing water", the word may also be applied to the inflow of the tide. Floods are an area of study of the discipline hydrol ...

avalanche An avalanche is a rapid flow of snow down a slope, such as a hill or mountain. Avalanches can be set off spontaneously, by such factors as increased precipitation or snowpack weakening, or by external means such as humans, animals, and earth ...

s and the like. Their causes are usually external (

exogenous In a variety of contexts, exogeny or exogeneity () is the fact of an action or object originating externally. It contrasts with endogeneity or endogeny, the fact of being influenced within a system. Economics In an economic model, an exogeno ...

) to the community—they are natural processes occurring (mostly) independently of the natural processes of the community (such as germination, growth, death, etc.). Such events can change vegetation structure and composition very quickly and for long time periods, and they can do so over large areas. Very few ecosystems are without some type of disturbance as a regular and recurring part of the long term

system A system is a group of Interaction, interacting or interrelated elements that act according to a set of rules to form a unified whole. A system, surrounded and influenced by its environment (systems), environment, is described by its boundaries, ...

dynamic.

Fire Fire is the rapid oxidation of a material (the fuel) in the exothermic chemical process of combustion, releasing heat, light, and various reaction Product (chemistry), products. At a certain point in the combustion reaction, called the ignition ...

and wind disturbances are particularly common throughout many vegetation types worldwide. Fire is particularly potent because of its ability to destroy not only living plants, but also the seeds, spores, and living

meristem The meristem is a type of tissue found in plants. It consists of undifferentiated cells (meristematic cells) capable of cell division. Cells in the meristem can develop into all the other tissues and organs that occur in plants. These cells conti ...

s representing the potential next generation, and because of fire's impact on fauna populations,

soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former te ...

characteristics and other ecosystem elements and processes (for further discussion of this topic see

fire ecology Fire ecology is a scientific discipline concerned with natural processes involving fire in an ecosystem and the ecological effects, the interactions between fire and the abiotic and biotic components of an ecosystem, and the role as an ecosystem p ...

). Temporal change at a slower pace is ubiquitous; it comprises the field of

ecological succession Ecological succession is the process of change in the species structure of an ecological community over time. The time scale can be decades (for example, after a wildfire) or more or less. Bacteria allows for the cycling of nutrients such as ca ...

. Succession is the relatively gradual change in structure and taxonomic composition that arises as the vegetation itself modifies various environmental variables over time, including light, water and

nutrient A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...

levels. These modifications change the suite of species most adapted to grow, survive and reproduce in an area, causing floristic changes. These floristic changes contribute to structural changes that are inherent in plant growth even in the absence of species changes (especially where plants have a large maximum size, i.e. trees), causing slow and broadly predictable changes in the vegetation. Succession can be interrupted at any time by disturbance, setting the system either back to a previous state, or off on another

trajectory A trajectory or flight path is the path that an object with mass in motion follows through space as a function of time. In classical mechanics, a trajectory is defined by Hamiltonian mechanics via canonical coordinates; hence, a complete traj ...

altogether. Because of this, successional processes may or may not lead to some static, final state. Moreover, accurately predicting the characteristics of such a state, even if it does arise, is not always possible. In short, vegetative communities are subject to many variables that together set limits on the predictability of future conditions.

Spatial dynamics

As a general rule, the larger an area under consideration, the more likely the vegetation will be heterogeneous across it. Two main factors are at work. First, the temporal dynamics of disturbance and succession are increasingly unlikely to be in synchrony across any area as the size of that area increases. That is, different areas will be at different developmental stages due to different local histories, particularly their times since last major disturbance. This fact interacts with inherent environmental variability (e.g. in soils, climate, topography, etc.), which is also a function of area. Environmental variability constrains the suite of species that can occupy a given area, and the two factors together interact to create a mosaic of vegetation conditions across the landscape. Only in

agricultural Agriculture or farming is the practice of cultivating Plant, plants and livestock. Agriculture was the key development in the rise of Sedentism, sedentary human civilization, whereby farming of Domestication, domesticated species created food ...

horticultural Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...

systems does vegetation ever approach perfect uniformity. In natural systems, there is always heterogeneity, although its scale and intensity will vary widely..

References

External links

Classification

Terrestrial Vegetation of the United States Volume I – The National Vegetation Classification System: Development, Status, and Applications
(PDF)
Federal Geographic Data Committee Vegetation Subcommittee

Vegetation Classification Standard
GDC-STD-005, June 1997(PDF)
Classifying Vegetation Condition: Vegetation Assets States and Transitions (VAST)

Climate diagrams

ClimateDiagrams.com
Provides climate diagrams for more than 3000 weather stations and for different climate periods from all over the world. Users can also create their own diagrams with their own data.

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