Variable Hawk
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The variable hawk (''Geranoaetus polyosoma'') is a polymorphic species of
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators h ...
in the family
Accipitridae The Accipitridae is one of the three families within the order Accipitriformes, and is a family of small to large birds with strongly hooked bills and variable morphology based on diet. They feed on a range of prey items from insects to medium-s ...
. It is widespread and often common in open habitats in western and southern South America, including the
Falkland Islands The Falkland Islands (; es, Islas Malvinas, link=no ) is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and about from Cape Dubouzet ...
. Its
taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
is disputed, with some splitting it into the widespread red-backed hawk (''G. polyosoma'') and the Puna hawk or Gurney's hawk (''G. poecilochrous'') of the central and north
Andean The Andes, Andes Mountains or Andean Mountains (; ) are the List of mountain ranges#Mountain ranges by length, longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range i ...
highlands, but the differences between the two are unclear. Most recent authorities have supported the lumping together of the two hawks although the issue still is controversial.South American Classification Committee (2007).
Merge Buteo poecilochrous into B. polyosoma.
'. Accessed 10-07-2009
South American Classification Committee (2009).

'. Accessed 10-07-2009
On the contrary, the rare
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
from the
Juan Fernández Islands The Juan Fernández Islands ( es, Archipiélago Juan Fernández) are a sparsely inhabited series of islands in the South Pacific Ocean reliant on tourism and fishing. Situated off the coast of Chile, they are composed of three main volcanic i ...
is relatively distinctive, and possibly worthy of species recognition as the Juan Fernández hawk (''B. exsul'').Jaramillo, A. Burke, P., & Beadle, D. (2003). Birds of Chile. Christopher Helm. In this article and most current accounts, the three races are regarded as subspecies.


Description

The name variable hawk is fully deserved, as both sexes occur in several morphs. Adults of all have a white tail with a contrasting black subterminal band and grey wings barred dark (in flight from below, the
remiges Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...
appear whitish with fine barring and a broad black tip). The remaining
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
varies from very dark grey to whitish, and some individuals have reddish-brown to the underparts. Females usually have a reddish-brown back, which males usually lack, although at least some males also have this. The taxon ''exsul'' from the Juan Fernández Islands is far less variable, being whitish below and grey above in adults of both sexes. At least 27 distinct adult plumages are known in this species, possibly the most of any
raptor Raptor or RAPTOR may refer to: Animals The word "raptor" refers to several groups of bird-like dinosaurs which primarily capture and subdue/kill prey with their talons. * Raptor (bird) or bird of prey, a bird that primarily hunts and feeds on ...
(although the widespread
red-tailed hawk The red-tailed hawk (''Buteo jamaicensis'') is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members with ...
&
Eurasian buzzard The common buzzard (''Buteo buteo'') is a medium-to-large bird of prey which has a large range. A member of the genus ''Buteo'', it is a member of the family Accipitridae. The species lives in most of Europe and extends its breeding range across ...
have also been noted for a widespread but more uniform spectrum of plumage variations) with no relationship to
morphometric Morphometrics (from Greek μορϕή ''morphe'', "shape, form", and -μετρία ''metria'', "measurement") or morphometry refers to the quantitative analysis of ''form'', a concept that encompasses size and shape. Morphometric analyses are co ...
variables and only minor geographic variation.


Size

Size is also variable in this confusing species. Length can range from 45 to 62 cm (18–25 in) and wingspan ranging from 113 to 151 cm (45–60 in). Weight can range from 800 to at least 1,800 grams (1.8-4 lbs). The Puna hawk subspecies generally considered to be at the larger end of the size spectrum and the red-backed hawk at the smaller (Juan Hernandez hawk being intermediate), but these distinctions are difficult at best in the field. Overall, however, this species rates as a large ''Buteo''.


Habitat

Variable hawks occupy open habitats at all elevations. The red-backed race inhabits the widest range of areas of the variable hawk races, including above
tree line The tree line is the edge of the habitat at which trees are capable of growing. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions (usually cold temperatures, extreme snowp ...
in mountains, Pacific coastal foothills,
Patagonian Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and g ...
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate grasslands, ...
s,
agricultural Agriculture or farming is the practice of cultivating Plant, plants and livestock. Agriculture was the key development in the rise of Sedentism, sedentary human civilization, whereby farming of Domestication, domesticated species created food ...
areas and edges of river galleries,
beech Beech (''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to temperate Europe, Asia, and North America. Recent classifications recognize 10 to 13 species in two distinct subgenera, ''Engleriana'' and ''Fagus''. The ''Engle ...
woods and humid premontane and lowland forests. Though often the most likely race to be found in lowlands, even the red-backed is less than common below an elevation of 500 m (up to 3,000 m). The Juan Fernandez race is found on the islands' volcanic slopes and barren grazed grasslands at all elevations. The Puna race are often a common element above
tree line The tree line is the edge of the habitat at which trees are capable of growing. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions (usually cold temperatures, extreme snowp ...
in
páramo Páramo () can refer to a variety of alpine tundra ecosystems located in the Andes Mountain Range, South America. Some ecologists describe the páramo broadly as "all high, tropical, montane vegetation above the continuous timberline". A narrower ...
and puna habitat, at higher elevations (5,000+ m) than almost any other raptor. Smaller numbers of the latter race may visit mountain scrub and stunted ''
Polylepis ''Polylepis'' is a genus comprising 28 recognised shrub and tree species, that are endemic to the mid- and high-elevation regions of the tropical Andes. This group is unique in the rose family in that it is predominantly wind-pollinated. They are ...
'' woodland at as low 900 m (but rarely below 2,900 m).


Behavior

They are most often seen soaring on warm thermals but may be seen on almost any type of raised perch (from sign posts to large trees). They prey on almost any small to medium-sized animals that can be caught, but smallish mammals comprise more than 90% of prey in some studies. The most commonly recorded prey includes
cavies Caviidae, the cavy family, is composed of rodents native to South America and includes the domestic guinea pig, wild cavies, and the largest living rodent, the capybara. They are found across South America in open areas from moist savanna to tho ...
,
tuco-tuco A tuco-tuco is a neotropical rodent in the family Ctenomyidae.Parada, A., G. D’Elia, C.J. Bidau, and E.P. Lessa. 2011. Species Groups and the Evolutionary Diversification of Tuco-Tucos, genus ''Ctenomys'' (Rodentia: Ctenomyidae). ''Journal of M ...
s,
rabbit Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit speci ...
s,
mice A mouse ( : mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...
and páramo rats.
Earthworm An earthworm is a terrestrial invertebrate that belongs to the phylum Annelida. They exhibit a tube-within-a-tube body plan; they are externally segmented with corresponding internal segmentation; and they usually have setae on all segments. Th ...
s,
weevil Weevils are beetles belonging to the Taxonomic rank, superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and Herbivore, herbivorous. Approximately 97,000 species of weevils are known. They b ...
s,
orthopteran Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grassho ...
s and other
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s are often taken. Birds are sometimes taken, including tired
petrel Petrels are tube-nosed seabirds in the bird order Procellariiformes. Description The common name does not indicate relationship beyond that point, as "petrels" occur in three of the four families within that group (all except the albatross f ...
s around
Juan Fernández Islands The Juan Fernández Islands ( es, Archipiélago Juan Fernández) are a sparsely inhabited series of islands in the South Pacific Ocean reliant on tourism and fishing. Situated off the coast of Chile, they are composed of three main volcanic i ...
. Other prey include other
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
s &
lagomorph The lagomorphs are the members of the taxonomic order Lagomorpha, of which there are two living families: the Leporidae (hares and rabbits) and the Ochotonidae (pikas). The name of the order is derived from the Ancient Greek ''lagos'' (λαγ ...
s,
frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...
s,
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s,
snake Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Ma ...
s, and
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
. The variable hawk hunts with prey being spotted while soaring from the air and pinned on the ground. Breeding is at various seasons and may be variable for all races. They build large stick nests on any elevated structure available, and sometimes breed cooperatively. One to three eggs are laid. The incubation period is 26 to 36 days. The nestlings fledge anywhere from 40 to 74 days. The larger-bodied, high-elevation hawks take longer to incubate and much longer to fledge than lower elevation hawks.


References

*"Raptors of the World" by Ferguson-Lees, Christie, Franklin, Mead & Burton. Houghton Mifflin (2001), . * * {{Taxonbar, from=Q1576679 variable hawk Birds of South America Birds of Chile Birds of Argentina Birds of Ecuador Birds of Peru Birds of Patagonia Birds of the Falkland Islands Páramo fauna variable hawk