Upeneus moluccensis
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''Upeneus moluccensis'', the goldband goatfish, golden-banded goatfish or Moluccan goatfish, is a species of Indo-Pacific goatfish from the
red mullet Red is the color at the long wavelength end of the visible spectrum of light, next to orange and opposite violet. It has a dominant wavelength of approximately 625–740 nanometres. It is a primary color in the RGB color model and a secondary ...
and
goatfish The goatfishes are perciform fish of the family Mullidae. The family is also sometimes referred to as the red mullets, which also refers more narrowly to the genus ''Mullus''. The family name and the English common name mullet derived from Lati ...
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
, the Mullidae. It is widespread in the warmer waters of the Indian and Pacific Oceans as far east as
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
and has colonised the eastern Mediterranean Sea from the Red Sea via the
Suez Canal The Suez Canal ( arz, قَنَاةُ ٱلسُّوَيْسِ, ') is an artificial sea-level waterway in Egypt, connecting the Mediterranean Sea to the Red Sea through the Isthmus of Suez and dividing Africa and Asia. The long canal is a popular ...
, making it a
Lessepsian migrant The Lessepsian migration (also called Erythrean invasion) is the migration of marine species across the Suez Canal, usually from the Red Sea to the Mediterranean Sea, and more rarely in the opposite direction. When the canal was completed in 18 ...
.


Description

''Upeneus moluccensis'' has an elongated body which has a sub-cylindrical anterior portion which becomes compressed towards the tail with two
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
s which are well separated, with the second dorsal fin directly above the
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
. The
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
is deeply forked. The chin bears a pair of barbels which do not extend past the margin of the
preoperculum This glossary of ichthyology is a list of definitions of terms and concepts used in ichthyology, the study of fishes. A B C ...
. The upper and lower jaws, the
palatine A palatine or palatinus (in Latin; plural ''palatini''; cf. derivative spellings below) is a high-level official attached to imperial or royal courts in Europe since Roman times.
and the
vomer The vomer (; lat, vomer, lit=ploughshare) is one of the unpaired facial bones of the skull. It is located in the midsagittal line, and articulates with the sphenoid, the ethmoid, the left and right palatine bones, and the left and right maxill ...
are covered in brush-like or villiform teeth. The back is pinkish-red in colour contrasting with the white belly, from which it is separated by single longitudinal yellow stripe which runs from the operculum to the caudal fin peduncle. The dorsal fins are yellow in colour with 3 parallel red bars while the pectoral fin is a similar colour but lacks markings and the
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two en ...
is colourless. The upper lobe of caudal fin is whitish and has 5 or 6 black diagonal bars while the lower lobe is unmarked except for the hind margin which is black as the second bar of the upper lobe extends along it. They can grow to 27 cm but are more usually 7–20 cm.Atlas of Exotic Fishes in the Mediterranean Sea (Upeneus moluccensis). 2nd Edition. 2021. 366p. CIESM Publishers, Paris, Monaco.https://ciesm.org/atlas/fishes_2nd_edition/Upeneus_moluccensis.pdf


Distribution

''Upeneus moluccensis'' has an
Indo-West Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
distribution, occurring from the
Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; T ...
to
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
and as far north as
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
, and south to northern Australia. It is the second most common member of its genus in New Caledonian waters. The first record of ''U. moluccensis'' in the Mediterranean was off
Palestine __NOTOC__ Palestine may refer to: * State of Palestine, a state in Western Asia * Palestine (region), a geographic region in Western Asia * Palestinian territories, territories occupied by Israel since 1967, namely the West Bank (including East ...
in 1947, it had reached the Mediterranean through the Suez Canal as a Lessepsian migrant, and has become common enough in the eastern Mediterranean to be an important species in fisheries.


Biology

''Upeneus moluccensis'' prefers sandy or muddy substrates where it searches for its food, mainly
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s using chemoreceptors on its barbels. As they grow larger fish form a greater part of their diet. In the Mediterranean they spawn from late July up to September, the spawn and larvae are planktonic, settling to a benthic habit at 4–5 cm in length and becoming sexually mature when they are about a year old, c. 10 cm in length. They form large schools which are normally quite fast swimmers, occasionally stopping to feed. It is normally found at depths of 10–80m on continental or insular shelf regions. The maximum lifespan for fish caught off Turkey was found to be 5 years for males and six years for females.


Fisheries

''Upeneus moluccensis'' is sold fresh in markets as well as being utilized for fish meal and valued for its roe. It is regarded as a "trash fish" by fishermen in the
South China Sea The South China Sea is a marginal sea of the Western Pacific Ocean. It is bounded in the north by the shores of South China (hence the name), in the west by the Indochinese Peninsula, in the east by the islands of Taiwan and northwestern Phil ...
because of its relatively small size. The annual catch of goatfish at the fishing harbour of
Visakhapatnam , image_alt = , image_caption = From top, left to right: Visakhapatnam aerial view, Vizag seaport, Simhachalam Temple, Aerial view of Rushikonda Beach, Beach road, Novotel, Novotel Visakhapatnam, INS Kursura (S20), INS ...
in India ranges between 2177 and 3463 tonnes, averaging 2859 tonnes, of which ''U. moluccensis'' makes up 18%. In the Mediterranean ''U. moluccensis'' is now an important commercial species for trawl fisheries.


References

{{Taxonbar, from=Q2482121 moluccensis Fish of the Indian Ocean Fish of the Pacific Ocean Fish of the Red Sea Fish described in 1855 Taxa named by Pieter Bleeker