Utahdactylus
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''Utahdactylus'' was a genus of
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
reptile from the Kimmeridgian-
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
-age
Upper Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
Morrison Formation The Morrison Formation is a distinctive sequence of Late Jurassic, Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandsto ...
of Utah, United States. Based on DM 002/CEUM 32588 (an incomplete skeleton described as including a fragment of the skull, a
cervical In anatomy, cervical is an adjective that has two meanings: # of or pertaining to any neck. # of or pertaining to the female cervix: i.e., the ''neck'' of the uterus. *Commonly used medical phrases involving the neck are **cervical collar **cervic ...
vertebra, three back vertebrae, and a
caudal Caudal may refer to: Anatomy * Caudal (anatomical term) (from Latin ''cauda''; tail), used to describe how close something is to the trailing end of an organism * Caudal artery, the portion of the dorsal aorta of a vertebrate that passes into the ...
vertebra, ribs, a scapula, coracoid, and limb bones), Czerkas and Mickelson (2002) identified it as a " rhamphorhynchoid" pterosaur. Bennett (2007) later concluded that it has no diagnostic features of the Pterosauria, and cannot be positively identified beyond being an indeterminate diapsid. More recent work on newly prepared material, however, seems to confirm once again that ''Utahdactylus'' was a pterosaur.


History

The genus was named and described in 2002 by Stephen Czerkas and Debra Mickelson. The type species is ''Utahdactylus kateae''. The genus name is derived from Utah and Greek ''daktylos'', "finger". The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
means "for Kate", referring to Kate Mickelson. The holotype consists of some disarticulated bone fragments preserved on several chalkstone blocks. It is housed in the Dinosaur Museum, run by Czerkas himself. The specimen was first described as a pterosaur, with a long tail and an estimated wingspan of 1.20 meters (3.94 feet). The authors considered it to be a "rhamphorhynchoid", i.e. a basal pterosaur, due to its long tail and large but not elongate cervical vertebrae, but without the typical groove in its forelimb bones. It was regarded as a "rhamphorhynchoid" based on an unprepared specimen in the most recent review of Morrison pterosaurs. In 2007, pterosaur specialist Chris Bennett published a redescription wherein he disagreed with Czerkas' and Mickelson's conclusions. He found several of the bone identifications and interpretations to be mistaken, such as the skull bone (interpreted here as just a bone fragment of unknown origin), elongate tail vertebra (the presumed elongated extensions were ribs), humerus (unknown), and the orientation of the bone described as a scapulacoracoid (the scapula and coracoid parts had been confused). He could not locate other bones seen as impressions, and found no evidence to suggest that the identifiable bones came from a pterosaur. In fact, he found the general quality of the bone texture to differ from that of pterosaur bones. He concluded by classifying it as Diapsida '' incertae sedis'', and a
dubious name In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...
, adding an exhortation not to name pterosaurs from material lacking unequivocal pterosaur characters. Bennett's conclusion was rejected by Czerkas & Ford (2018), who affirmed the validity of initial interpretation of ''Utahdactylus'' as a pterosaur. The authors reported that the cervical of the holotype specimen was completely prepared out of the matrix along with the scapulacoracoid, a sacrum, and part of a mandible. The study of the glenoid indicated that it had a pterosaur saddle-shaped articulation for the
humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...
. In addition, the authors reported a parallel alignment of the
dentaries In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
, indicating that they formed a
mandibular symphysis In human anatomy, the facial skeleton of the skull the external surface of the mandible is marked in the median line by a faint ridge, indicating the mandibular symphysis (Latin: ''symphysis menti'') or line of junction where the two lateral halves ...
. Czerkas & Ford interpreted the presence of the elongate mandibular symphysis and spoon-like expansion of the anterior end of the jaws as aligning ''Utahdactylus'' with pterodactyloids, and specifically within Ctenochasmatidae. The authors interpreted ''Utahdactylus'' as the first known gnathosaur from North America.


See also

* List of pterosaur genera * Timeline of pterosaur research


References


External links


''Utahdactylus''
in The Pterosauria {{Taxonbar, from=Q1014541, from2=Q18602545 Pterosaurs Nomina dubia Late Jurassic pterosaurs of North America Morrison fauna Fossil taxa described in 2002