The Ussuri dholeHeptner, V. G. & Naumov, N. P. (1998).
Mammals of the Soviet Union Vol. II Part 1a, SIRENIA AND CARNIVORA (Sea cows; Wolves and Bears)
', Science Publishers, Inc. USA., pp. 566–86, (''Cuon alpinus alpinus''), also known as the Indian dhole, Eastern Asiatic dhole, Chinese dhole or southern dhole, is the

nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

of the

dhole The dhole (''Cuon alpinus''; ) is a canid native to Central, South, East and Southeast Asia. Other English names for the species include Asian wild dog, Asiatic wild dog, Indian wild dog, whistling dog, red dog, red wolf, and mountain wolf. It ...

native to

East Asia East Asia is the eastern region of Asia, which is defined in both geographical and ethno-cultural terms. The modern states of East Asia include China, Japan, Mongolia, North Korea, South Korea, and Taiwan. China, North Korea, South Korea and ...

. The Ussuri dhole is also native to

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

, however it is probably extinct in most of its ranges in China, as well as in

Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...

and the

Russian Far East The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in Northeast Asia. It is the easternmost part of Russia and the Asian continent; and is admini ...

Physical descriptions

The Ussuri dhole is the largest subspecies. It has a bright red coat and a narrow skull. Like the Tian Shan dhole, the Ussuri dhole has a woolly winter coat, white underfur and larger mane during the cold season. On the other hand, the summer coat is coarser and leaner. The dholes from southern Asia has darker coarse and leaner fur which is mostly found in the

Sumatran dhole The dhole (''Cuon alpinus''; ) is a canid native to Central, South, East and Southeast Asia. Other English names for the species include Asian wild dog, Asiatic wild dog, Indian wild dog, whistling dog, red dog, red wolf, and mountain wolf. It ...

, with the red underfur nearly visible.

Habitat and distribution

The Ussuri dhole lives in

forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...

s, plains,

grasslands A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natural ...

savannah A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the Canopy (forest), canopy does not close. The open canopy allows sufficient light to rea ...

steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate grasslands, ...

s and

alpine tundra Alpine tundra is a type of natural region or biome that does not contain trees because it is at high elevation, with an associated alpine climate, harsh climate. As the latitude of a location approaches the poles, the threshold elevation for alp ...

. It is the most widespread subspecies, ranging from

South South is one of the cardinal directions or Points of the compass, compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Pro ...

Northeast Asia Northeast Asia or Northeastern Asia is a geographical subregion of Asia; its northeastern landmass and islands are bounded by the Pacific Ocean. The term Northeast Asia was popularized during the 1930s by American historian and political scient ...

. It occurs in

India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...

Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...

, China,

Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...

Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...

and

Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...

. It was believed to be extinct in Mongolia,

Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...

and

Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic o ...

, though it is disputed that dholes persist in those countries.

Hunting and diet

The Indian wild dogs feed on small and larger herbivorous mammals such as

chital The chital or cheetal (''Axis axis''; ), also known as the spotted deer, chital deer, and axis deer, is a deer species native to the Indian subcontinent. It was first described and given a binomial name by German naturalist Johann Christian Po ...

muntjac Muntjacs ( ), also known as the barking deer or rib-faced deer, (URL is Google Books) are small deer of the genus ''Muntiacus'' native to South Asia and Southeast Asia. Muntjacs are thought to have begun appearing 15–35 million years ago, ...

, sambar,

barasingha The barasingha (''Rucervus duvaucelii''), also known as the swamp deer, is a deer species distributed in the Indian subcontinent. Populations in northern and central India are fragmented, and two isolated populations occur in southwestern Nepal. ...

chousingha The four-horned antelope (''Tetracerus quadricornis''), or ''chousingha'', is a small antelope found in India and Nepal. Its four Horn (anatomy), horns distinguish it from most other bovids, which have two horns (with a few exceptions, such as t ...

chinkara The chinkara (''Gazella bennettii''), also known as the Indian gazelle, is a gazelle species native to Iran, Afghanistan, Pakistan and India. Taxonomy The following six subspecies are considered valid: * Deccan chinkara (''G. b. bennettii'') ...

blackbuck The blackbuck (''Antilope cervicapra''), also known as the Indian antelope, is an antelope native to India and Nepal. It inhabits grassy plains and lightly forested areas with perennial water sources. It stands up to high at the shoulder. Ma ...

hog deer ''Axis'' is a genus of deer occurring in South and Southeast Asia. As presently defined by most authorities, four species are placed in the genus. Three of the four species are called hog deer. The genus name is a word mentioned in Pliny the Eld ...

wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The species is ...

water buffalo The water buffalo (''Bubalus bubalis''), also called the domestic water buffalo or Asian water buffalo, is a large bovid originating in the Indian subcontinent and Southeast Asia. Today, it is also found in Europe, Australia, North America, So ...

nilgai The nilgai (''Boselaphus tragocamelus'') (, literally meaning "blue cow") is the largest Asian antelope and is ubiquitous across the northern Indian subcontinent. It is the sole member of the genus ''Boselaphus'' and was described by Peter Sim ...

and

gaur The gaur (''Bos gaurus''; ), also known as the Indian bison, is a bovine native to South Asia and Southeast Asia, and has been listed as Vulnerable on the IUCN Red List since 1986. The global population was estimated at a maximum of 21,000 m ...

. On few occasions, Ussuri dholes also hunt

elephant Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae an ...

and

rhino A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species o ...

calves, and may also feed on

onager The onager (; ''Equus hemionus'' ), A new species called the kiang (''E. kiang''), a Tibetan relative, was previously considered to be a subspecies of the onager as ''E. hemionus kiang'', but recent molecular studies indicate it to be a distinct ...

s and

kiang The kiang (''Equus kiang'') is the largest of the '' Asinus'' subgenus. It is native to the Tibetan Plateau, where it inhabits montane and alpine grasslands. Its current range is restricted to the plains of the Tibetan plateau; Ladakh; and nort ...

Threats and enemies

Dholes are listed as an

endangered species An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...

due to low densities. However, threats such as

poaching Poaching has been defined as the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set a ...

, illegal hunting and the

fur trade The fur trade is a worldwide industry dealing in the acquisition and sale of animal fur. Since the establishment of a world fur market in the early modern period, furs of boreal, polar and cold temperate mammalian animals have been the mos ...

no longer pose significant threats to dholes.Durbin, L.S., Venkataraman, A., Hedges, S. & Duckworth, W. (2004)
Dhole ''Cuon alpinus'' (Pallas 1811)
in Sillero-Zubiri, C., Hoffmann, M. & Macdonald, D.W. (eds.) ''Canids: Foxes, Wolverhampton Wanderers F.C., Jackals and Dogs: Status Survey and Conservation Action Plan''. IUCN/SSC Canid Specialist Group. Gland, Switzerland and Cambridge, UK. x + pp. 210–219 The species is highly protected in many countries, such as in

Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand t ...

. The dholes in certain regions are mostly threatened by lack of prey and habitat loss. Dholes are also vulnerable to diseases from areas where they share the same habitats with other

canid Canidae (; from Latin, '' canis'', " dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found withi ...

s like

wolves The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...

and Eurasian golden jackals. Ussuri dholes may also be threatened by fellow apex predators such as

tiger The tiger (''Panthera tigris'') is the largest living cat species and a member of the genus '' Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily preys on u ...

leopard The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, a ...

, striped hyenas and

bear Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the Nor ...

s, and formerly by

Asiatic lion The Asiatic lion is a population of ''Panthera leo leo'' that today survives in the wild only in India. Since the turn of the 20th century, its range has been restricted to Gir National Park and the surrounding areas in the Indian state of Gujarat ...

s and

Asiatic cheetah The Asiatic cheetah (''Acinonyx jubatus venaticus'') is a critically endangered cheetah subspecies currently only surviving in Iran. It once occurred from the Arabian Peninsula and the Near East to the Caspian region, Transcaucasus, Kyzylkum D ...

s.Pocock, R. I. (1939)
''The Fauna of British India, including Ceylon and Burma. Mammalia. – Volume 1''
Taylor and Francis Ltd., London. Pp. 1–565.

References

{{Taxonbar, from=Q20907534 Dhole Mammals of East Asia Fauna of South Asia Mammals of India Mammals of Nepal Mammals of Korea Mammals of Thailand Mammals of Mongolia Mammals of China Mammals of Russia