The red-faced cormorant (''Urile urile''), red-faced shag or violet shag, is a

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

species of the family

Phalacrocoracidae Phalacrocoracidae is a family of approximately 40 species of aquatic birds commonly known as cormorants and shags. Several different classifications of the family have been proposed, but in 2021 the IOC adopted a consensus taxonomy of seven gen ...

. Its range spans from the eastern tip of

Hokkaidō is Japan's second largest island and comprises the largest and northernmost prefecture, making up its own region. The Tsugaru Strait separates Hokkaidō from Honshu; the two islands are connected by the undersea railway Seikan Tunnel. The la ...

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

, northern korean peninsula, via the

Kuril Islands The Kuril Islands or Kurile Islands (; rus, Кури́льские острова́, r=Kuril'skiye ostrova, p=kʊˈrʲilʲskʲɪjə ɐstrɐˈva; Japanese: or ) are a volcanic archipelago currently administered as part of Sakhalin Oblast in the ...

, the southern tip of the

Kamchatka Peninsula The Kamchatka Peninsula (russian: полуостров Камчатка, Poluostrov Kamchatka, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and we ...

and the

Aleutian Arc The Aleutian Arc is a large volcanic arc in the U.S. state of Alaska. It consists of a number of active and dormant volcanoes that have formed as a result of subduction along the Aleutian Trench. Although taking its name from the Aleutian Islands ...

to the

Alaska Peninsula The Alaska Peninsula (also called Aleut Peninsula or Aleutian Peninsula, ale, Alasxix̂; Sugpiaq: ''Aluuwiq'', ''Al'uwiq'') is a peninsula extending about to the southwest from the mainland of Alaska and ending in the Aleutian Islands. The ...

and

Gulf of Alaska The Gulf of Alaska (Tlingit: ''Yéil T'ooch’'') is an arm of the Pacific Ocean defined by the curve of the southern coast of Alaska, stretching from the Alaska Peninsula and Kodiak Island in the west to the Alexander Archipelago in the east, ...

Taxonomy

The red-faced cormorant was formally described in 1789 by the German naturalist

Johann Friedrich Gmelin , fields = , workplaces = University of GöttingenUniversity of Tübingen , alma_mater = University of Tübingen , doctoral_advisor = Philipp Friedrich GmelinFerdinand Christoph Oetinger , academic_advisors = , doctora ...

in his revised and expanded edition of

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...

's ''

Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...

''. He placed it in the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

'' Pelecanus'' and coined the

binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

''Pelecanus urile''. Gmelin based his description on the "red faced shag" of John Latham and the "red faced corvarant" of

Thomas Pennant Thomas Pennant (14 June Old Style, OS 172616 December 1798) was a Welsh natural history, naturalist, traveller, writer and antiquarian. He was born and lived his whole life at his family estate, Downing Hall near Whitford, Flintshire, in Wales ...

. The authors cited different translations of an account of

Georg Wilhelm Steller Georg Wilhelm Steller (10 March 1709 – 14 November 1746) was a German botanist, zoologist, physician and explorer, who worked in Russia and is considered a pioneer of Alaskan natural history.Evans, Howard Ensign. Edward Osborne Wilson (col.) ...

's exploration of the

. The red-faced cormorant was formerly classified in the genus ''

Phalacrocorax ''Phalacrocorax'' is a genus of fish-eating birds in the cormorant family Phalacrocoracidae. Members of this genus are also known as the Old World cormorants. Taxonomy The genus ''Phalacrocorax'' was introduced by the French zoologist Mathurin ...

''; it was moved to the resurrected genus ''

Urile ''Urile'' is a genus of birds in the family Phalacrocoracidae, commonly known as North Pacific cormorants. It contains 3 extant and 1 recently extinct species, all of which are or were found in the North Pacific Ocean. Members of this genus were ...

'' based on a

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

study published in 2014. The genus had been introduced in 1856 by the French naturalist

Charles Lucien Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857), was a French naturalist and ornithologist. Lucien and his wife had twelve children, including Cardinal Lucien Bonaparte. Life and career ...

. The epithet ''urile'' is possibly derived from the Russian vernacular name of this species, referring to the

where this species is now very rare. The species is

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...

: no

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

are recognised. Within the genus ''Urile'' the red-faced cormorant is closely related to the

pelagic cormorant The pelagic cormorant (''Urile pelagicus''), also known as Baird's cormorant or violet-green cormorant, is a small member of the cormorant family Phalacrocoracidae. Analogous to other smallish cormorants, it is also called the pelagic shag occasi ...

(''Urile pelagicus'').

Description

The adult bird has glossy plumage that is a deep greenish blue in color, becoming purplish or bronze on the back and sides. In breeding condition it has a double crest, and white plumes on the flanks, neck and rump, and the bare facial skin of the lores and around the eyes is a bright orange or red, giving the bird its name; although the coloration is less vivid outside the breeding season, the red facial skin is enough to distinguish it from the otherwise rather similar pelagic cormorant. Its legs and feet are brownish black. Its wings range from 25 to 29 cm in extent, with females having on average about 5 cm shorter wings. Adults weigh between 1.5 and 2.3 kg, with females averaging 350 g less than males.

Behaviour

Breeding

Where it nests alongside the pelagic cormorant, the red-faced cormorant generally breeds the more successfully of the two species, and it is currently increasing in numbers, at least in the easterly parts of its range. It is however listed as being of conservation concern, partly because relatively little is so far known about it.

Food and feeding

Analysis of stomach contents suggests that the red-faced cormorant is mainly a bottom feeder, taking cottids especially. Adults have few predators, though river otters may attempt to take them, as will

corvid Corvidae is a Cosmopolitan distribution, cosmopolitan family (biology), family of oscine passerine birds that contains the crows, ravens, rook (bird), rooks, jackdaws, jays, magpies, treepies, choughs, and nutcracker (bird), nutcrackers. In coll ...

s of various species,

bald eagle The bald eagle (''Haliaeetus leucocephalus'') is a bird of prey found in North America. A sea eagle, it has two known subspecies and forms a species pair with the white-tailed eagle (''Haliaeetus albicilla''), which occupies the same niche as ...

s and

golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of p ...

s. Gulls and corvids are common predators on eggs and chicks.

References

{{Taxonbar, from=Q926664

red-faced cormorant The red-faced cormorant (''Urile urile''), red-faced shag or violet shag, is a bird species of the family Phalacrocoracidae. Its range spans from the eastern tip of Hokkaidō in Japan, northern korean peninsula, via the Kuril Islands, the sou ...

Birds of the Aleutian Islands

Red-faced cormorant The red-faced cormorant (''Urile urile''), red-faced shag or violet shag, is a bird species of the family Phalacrocoracidae. Its range spans from the eastern tip of Hokkaidō in Japan, northern korean peninsula, via the Kuril Islands, the sou ...