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''Toona sureni'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
in the mahogany family. It is native to
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical Geography (from Greek: , ''geographia''. Combination of Greek words ‘Geo’ (The Earth) and ‘Graphien’ (to describe), literally "earth descr ...
,
Indochina Mainland Southeast Asia, also known as the Indochinese Peninsula or Indochina, is the continental portion of Southeast Asia. It lies east of the Indian subcontinent and south of Mainland China and is bordered by the Indian Ocean to the west an ...
,
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms, and also a phytogeographical floristic region in the Paleotropical Kingdom. It has been given different definitions. Th ...
, China, and
Papua New Guinea Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
. It is commonly known as the suren toon, surian, limpaga, iron redwood or the red cedar (a name also shared with various other trees). It is also known as the Indonesian mahogany or the Vietnamese mahogany. The species is a valuable timber tree.


Taxonomy

''Toona sureni'' belongs to the toon
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
'' Toona'' of the mahogany
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Meliaceae Meliaceae, the mahogany family, is a flowering plant family of mostly trees and shrubs (and a few herbaceous plants, mangroves) in the order Sapindales. They are characterised by alternate, usually pinnate leaves without stipules, and by syncar ...
. It was first described by the German-Dutch botanist
Carl Ludwig Blume Charles Ludwig de Blume or Karl Ludwig von Blume (9 June 1796, Braunschweig – 3 February 1862, Leiden) was a German- Dutch botanist. He was born at Braunschweig in Germany, but studied at Leiden University and spent his professional life wor ...
in 1823 as ''Swietenia sureni''. It was transferred to the genus ''Toona'' in 1917 by the American botanist
Elmer Drew Merrill Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through t ...
.


Description

''Toona sureni'' is a medium-sized to large tree, reaching a maximum height of around and a diameter of . The bole possesses
buttress root Buttress roots also known as plank roots are large, wide roots on all sides of a shallowly rooted tree. Typically, they are found in nutrient-poor tropical forest soils that may not be very deep. They prevent the tree from falling over (hence t ...
s up to a height of and remains unbranched and straight up to a height of . The bark is fibrous and flaky in texture with numerous vertical fissures. It is pale brown to whitish or grayish brown in color. When cut, the bark and sapwood produce a pleasant odor reminiscent of cinnamon. The
sap Sap is a fluid transported in xylem cells (vessel elements or tracheids) or phloem sieve tube elements of a plant. These cells transport water and nutrients throughout the plant. Sap is distinct from latex, resin, or cell sap; it is a separ ...
is colorless and does not change in color after being exposed to air. The sapwood of ''Toona sureni'' is white to pinkish or light red in color, while the
heartwood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin t ...
is light red or brown. The leaves are
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
and large, with a length of about . They are arranged spirally, usually clustered at the ends of twigs. The leaflets are lanceolate to ovate-lanceolate in shape. They are arranged somewhat opposite each other, usually in pairs of 6 to 9, with a maximum of 12 pairs. The upper and lower surfaces of the leaflet midribs are characteristically hairy (pilose). The
inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...
are terminal, occurring at the tips of branchlets. They are paniculate and pendant, reaching a length of around . The individual flowers are small (around in length), and are sweetly fragrant. Flowers are individually unisexual, though both male and female flowers occur in the same plant. The fruits are leathery capsules, around long and brown in color. Each contains more than 100 seeds. The seeds are narrow and usually in length, with a maximum length of ; and in diameter. They are winged at both ends (subequal).


Ecology

''Toona sureni'' is
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...
, shedding their leaves during the dry season (usually February to March or September to October). They produce flowers and fruits twice each year (usually during December to February and April to September).


Distribution and habitat

''Toona sureni'' is native to
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical Geography (from Greek: , ''geographia''. Combination of Greek words ‘Geo’ (The Earth) and ‘Graphien’ (to describe), literally "earth descr ...
(
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
,
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainou ...
, and
Nepal Nepal (; ne, :ne:नेपाल, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in S ...
);
Indochina Mainland Southeast Asia, also known as the Indochinese Peninsula or Indochina, is the continental portion of Southeast Asia. It lies east of the Indian subcontinent and south of Mainland China and is bordered by the Indian Ocean to the west an ...
(
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
, Laos,
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailan ...
,
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
, and
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
); China (
Guizhou Guizhou (; formerly Kweichow) is a landlocked province in the southwest region of the People's Republic of China. Its capital and largest city is Guiyang, in the center of the province. Guizhou borders the autonomous region of Guangxi to the ...
,
Hainan Hainan (, ; ) is the smallest and southernmost province of the People's Republic of China (PRC), consisting of various islands in the South China Sea. , the largest and most populous island in China,The island of Taiwan, which is slightly l ...
,
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
, and
Yunnan Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the C ...
);
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms, and also a phytogeographical floristic region in the Paleotropical Kingdom. It has been given different definitions. Th ...
(
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
, and the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
); and
Papua New Guinea Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
. They are usually found in
primary forest An old-growth forestalso termed primary forest, virgin forest, late seral forest, primeval forest, or first-growth forestis a forest that has attained great age without significant disturbance, and thereby exhibits unique ecological feature ...
s on open hillsides, slopes, ravines, and riverbanks at an altitude of asl. They may sometimes also be found in secondary semi-
evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, Live oak, acacias, magnolia, and banksia in more temperate zo ...
s.


Names

The tree is commonly known in English as the "suren toon" (or "suren toona"), "surian", "limpaga", "iron redwood" or the "red cedar". It is also sometimes known as the "Indonesian mahogany" or the "Vietnamese mahogany", though it is not a true mahogany (genus ''
Swietenia ''Swietenia'' is a genus of trees in the chinaberry family, Meliaceae. It occurs natively in the Neotropics, from southern Florida, the Caribbean, Mexico and Central America south to Bolivia. The genus is named for Dutch-Austrian physician Ge ...
''). Local names include ''suren'' in Indonesia, ''ye tama'' in Burma, ''danupra'' in the Philippines, ''surian'' (สุเหรียน) in Thailand, ''surian wangi'' in Malaysia, and ''zi chun'' (紫椿) in China.


Uses

''Toona sureni'', like other members of the mahogany family, are valuable timber trees. They are a source of high quality commercial hardwoods used for high-end furniture work, interior finishing, decorative paneling, musical instruments such as Djembe, and other wood crafts. The bark extract is also used in
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
as an
astringent An astringent (sometimes called adstringent) is a chemical that shrinks or constricts body tissues. The word derives from the Latin ''adstringere'', which means "to bind fast". Calamine lotion, witch hazel, and yerba mansa, a Californian pla ...
,
purgative Laxatives, purgatives, or aperients are substances that loosen stools and increase bowel movements. They are used to treat and prevent constipation. Laxatives vary as to how they work and the side effects they may have. Certain stimulant, lubri ...
, antirheumatic, and for treating gastrointestinal ailments like
diarrhea Diarrhea, also spelled diarrhoea, is the condition of having at least three loose, liquid, or watery bowel movements each day. It often lasts for a few days and can result in dehydration due to fluid loss. Signs of dehydration often begin w ...
and
dysentery Dysentery (UK pronunciation: , US: ), historically known as the bloody flux, is a type of gastroenteritis that results in bloody diarrhea. Other symptoms may include fever, abdominal pain, and a feeling of incomplete defecation. Complications ...
. The leaf extracts are used as
antibacterial An antibiotic is a type of antimicrobial substance active against bacteria. It is the most important type of antibacterial agent for fighting bacterial infections, and antibiotic medications are widely used in the treatment and prevention ...
poultices. They are also planted as ornamentals and
shade tree A shade tree is a large tree whose primary role is to provide shade in the surrounding environment due to its spreading canopy and crown, where it may give shelter from sunlight in the heat of the summer for people who seek recreational needs i ...
s, as well as being used for
intercropping Intercropping is a multiple cropping practice that involves growing two or more crops in proximity. In other words, intercropping is the cultivation of two or more crops simultaneously on the same field. The most common goal of intercropping is ...
.


References

{{Taxonbar, from=Q15304020 sureni Plants described in 1823 Trees of Cambodia Trees of Nepal Trees of Vietnam Flora of Indo-China