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The thrushes are a
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
, Turdidae, with a worldwide distribution. The family was once much larger before biologists reclassified the former subfamily Saxicolinae, which includes the chats and European robins, as
Old World flycatcher The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Norther ...
s. Thrushes are small to medium-sized ground living birds that feed on insects, other invertebrates and fruit. Some unrelated species around the world have been named after thrushes due to their similarity to birds in this family.


Characteristics

Thrushes are plump, soft-plumaged, small to medium-sized birds, inhabiting wooded areas, and often feeding on the ground. The smallest thrush may be the
forest rock thrush The forest rock thrush (''Monticola sharpei'') is a songbird in the family Muscicapidae, formerly placed in the Turdidae together with the other chats. It now includes Benson's rock thrush and Amber Mountain rock thrush as subspecies. It is ...
, at and . However, the shortwings, which have ambiguous alliances with both thrushes and
Old World flycatcher The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Norther ...
s, can be even smaller. The
lesser shortwing The lesser shortwing (''Brachypteryx leucophris'') is a species of chat. This species is now classified in the family Muscicapidae. It is found in south-eastern Asia, Sumatra, Java and the Lesser Sundas. Its natural habitat is subtropical or t ...
averages . The largest thrush is the
Great thrush The great thrush (''Turdus fuscater'') is a species of bird in the family Turdidae. It is found in Bolivia, Colombia, Ecuador, Peru, and Venezuela. It is considered as the largest thrush in South America. The great thrush's size distinguishes it ...
at and , though the commonly recognized Blue whistling-thrush is an
Old world flycatcher The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Norther ...
. The
Amami thrush The Amami thrush (''Zoothera major'') is a member of the Thrush (bird), thrush family Turdidae. It is endemic to the islands of Amami Ōshima and Kakeroma island in the northern Ryūkyū Islands, Nansei Islands of Japan. Description This large, ...
might, however, grow larger than the
Great thrush The great thrush (''Turdus fuscater'') is a species of bird in the family Turdidae. It is found in Bolivia, Colombia, Ecuador, Peru, and Venezuela. It is considered as the largest thrush in South America. The great thrush's size distinguishes it ...
. Most species are grey or brown in colour, often with speckled underparts. They are insectivorous, but most species also eat worms,
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial gastropod mollusks that have shells (those without shells are known as ...
s, and fruit (usually
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
). Many species are permanently resident in warm climates, while others migrate to higher latitudes during summer, often over considerable distances. Thrushes build cup-shaped
nests A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materi ...
, sometimes lining them with mud. They lay two to five speckled eggs, sometimes laying two or more
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts). ...
es per year. Both parents help in raising the young. In almost all cases, the nest is placed on a branch; the only exceptions are the three species of
bluebird The bluebirds are a North American group of medium-sized, mostly insectivorous or omnivorous birds in the order of Passerines in the genus ''Sialia'' of the thrush family (Turdidae). Bluebirds are one of the few thrush genera in the Americas. ...
, which nest in holes.


Ecology

Turdidae species spread the seeds of plants, contributing to the dispersal of many species and the recovery of ecosystems. Plants have limited
seed dispersal In Spermatophyte plants, seed dispersal is the movement, spread or transport of seeds away from the parent plant. Plants have limited mobility and rely upon a variety of dispersal vectors to transport their seeds, including both abiotic vectors, ...
mobility away from the parent plant and consequently rely upon a variety of
dispersal vector A dispersal vector is an agent of biological dispersal that moves a dispersal unit, or organism, away from its birth population to another location or population in which the individual will reproduce. These dispersal units can range from pollen t ...
s to transport their propagules, including both
abiotic In biology and ecology, abiotic components or abiotic factors are non-living chemical and physical parts of the environment that affect living organisms and the functioning of ecosystems. Abiotic factors and the phenomena associated with them under ...
and biotic vectors. Seeds can be dispersed away from the parent plant individually or collectively, as well as dispersed in both space and time. Many bats and birds rely heavily on fruits for their diet, including birds in the families Cotingidae,
Columbidae Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
,
Trogonidae The trogons and quetzals are birds in the order Trogoniformes which contains only one family, the Trogonidae. The family Trogonidae contains 46 species in seven genera. The fossil record of the trogons dates back 49 million years to the Early E ...
, Turdidae, and
Ramphastidae Toucans (, ) are members of the Neotropical near passerine bird family Ramphastidae. The Ramphastidae are most closely related to the American barbets. They are brightly marked and have large, often colorful bills. The family includes five g ...
. While eating fruit, these animals swallow seeds and then later regurgitate them or pass them in their faeces. Such ornithochory has been a major mechanism of seed dispersal across ocean barriers. Other seeds may stick to the feet or feathers of birds, and in this way may travel long distances. Seeds of grasses, spores of algae, and the eggs of molluscs and other invertebrates commonly establish in remote areas after long journeys of this sort. The Turdidae have a great ecological importance because some populations migrate long distances and disperse the seeds of endangered plant species at new sites, helping to eliminate inbreeding and increasing the genetic diversity of local flora.


Taxonomy

The family Turdidae was introduced (as Turdinia) by the French
polymath A polymath ( el, πολυμαθής, , "having learned much"; la, homo universalis, "universal human") is an individual whose knowledge spans a substantial number of subjects, known to draw on complex bodies of knowledge to solve specific pro ...
Constantine Samuel Rafinesque Constantine Samuel Rafinesque-Schmaltz (; October 22, 1783September 18, 1840) was a French 19th-century polymath born near Constantinople in the Ottoman Empire and self-educated in France. He traveled as a young man in the United States, ultimat ...
in 1815. The taxonomic treatment of this large family has varied significantly in recent years. Traditionally, the Turdidae included the small Old World species, like the
nightingale The common nightingale, rufous nightingale or simply nightingale (''Luscinia megarhynchos''), is a small passerine bird best known for its powerful and beautiful song. It was formerly classed as a member of the thrush family Turdidae, but is no ...
and
European robin The European robin (''Erithacus rubecula''), known simply as the robin or robin redbreast in Great Britain & Ireland, is a small insectivorous passerine bird that belongs to the chat subfamily of the Old World flycatcher family. About in len ...
in the subfamily Saxicolinae, but most authorities now place this group in the
Old World flycatcher The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Norther ...
family
Muscicapidae The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Norther ...
.
Molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
analysis has shown that the family Turdidae is a member of the
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Muscicapoidea and is
sister A sister is a woman or a girl who shares one or more parents with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a familial relationship, it is sometimes used endearingly to refer to ...
to the family
Muscicapidae The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Norther ...
. The two families diverged in the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
around 17 million years ago. The family formerly included more species. At the time of the publication of the third edition of
Howard and Moore Complete Checklist of the Birds of the World The ''Howard and Moore Complete Checklist of the Birds of the World'' is a book by Richard Howard and Alick Moore which presents a list of the bird species of the world. It was the first single-volume world bird list to include subspecies names, ...
in 2003, the genera ''
Myophonus The whistling thrushes comprise a genus ''Myophonus'' (''Myiophoneus'') of the Old World flycatcher family Muscicapidae. They are all medium-sized mostly insectivorous or omnivorous birds. They are all brightly coloured species found in India an ...
'', '' Alethe'', ''
Brachypteryx ''Brachypteryx'' is a genus of passerine birds containing six species known as shortwings, that occurs in southeast Asia. Shortwings are small birds with long legs, finely pointed bills, short tails and short rounded wings. They are shy elusive ...
'' and '' Heinrichia'' were included in Turdidae. Subsequent
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
studies have shown that the species in these four genera are more closely related to species in the family
Muscicapidae The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Norther ...
. As a consequence, these four genera are now placed in Muscicapidae. In contrast, the genus ''
Cochoa The cochoas (from ''cocho'', Nepali for '' Cochoa purpurea'') are medium-sized frugivorous, insectivorous and molluscivorous birds in the genus ''Cochoa''. Their bright contrasting plumage patterns, sexual dimorphism and feeding habits made thei ...
'' which had previously been placed in Muscicapidae was shown to belong in Turdidae.


Genera

The family contains 174 species which are divided into 18 genera: * '' Grandala'' – grandala * ''
Sialia The bluebirds are a North American group of medium-sized, mostly insectivorous or omnivorous birds in the order of Passerines in the genus ''Sialia'' of the thrush family (Turdidae). Bluebirds are one of the few thrush genera in the Americas. ...
'' – bluebirds (3 species) * ''
Stizorhina The rufous thrushes, also known as flycatcher-thrushes, are medium-sized insectivorous birds in the genus ''Stizorhina'' of the thrush family Turdidae. These are African forest dwelling species. They are sometimes placed in the genus ''Neocossyp ...
'' – rufous thrushes (2 species) * ''
Neocossyphus The ant thrushes are medium-sized insectivorous birds in the genus ''Neocossyphus'' of the thrush family Turdidae. These are African forest dwelling species In biology, a species is the basic unit of classification and a taxonomic rank o ...
'' – ant thrushes (2 species) * ''
Pinarornis The boulder chat (''Pinarornis plumosus'') is a species of bird in the thrush family Turdidae native to dry woodland and savanna in Southern Africa. It is the only species in the monotypic genus ''Pinarornis''. Taxonomy The boulder chat was f ...
'' – boulder chat * ''
Myadestes ''Myadestes'' is a genus of solitaires, medium-sized mostly insectivorous birds in the thrush family, Turdidae. They are found in the Americas and Hawaii, where several island species have become extinct. Species in taxonomic order * ''Mya ...
'' – solitaires (12 species, including one recently extinct) * '' Chlamydochaera'' – fruithunter * ''
Cochoa The cochoas (from ''cocho'', Nepali for '' Cochoa purpurea'') are medium-sized frugivorous, insectivorous and molluscivorous birds in the genus ''Cochoa''. Their bright contrasting plumage patterns, sexual dimorphism and feeding habits made thei ...
'' – cochoas (4 species) * '' Ixoreus'' – varied thrush * '' Ridgwayia'' – Aztec thrush * ''
Cichlopsis The rufous-brown solitaire (''Cichlopsis leucogenys'') is a species of bird in the thrush family Turdidae. It is found in Brazil, Colombia, Ecuador, Guyana, Peru, Suriname, and Venezuela. Its natural habitats are subtropical or tropical moist ...
'' – rufous-brown solitaire * ''
Entomodestes ''Entomodestes '' is a small genus of birds in the thrush family. They are found in humid Andean The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland a ...
'' – solitaires (2 species) * '' Hylocichla'' – wood thrush * ''
Catharus The genus ''Catharus'' is an evolutionary clade of forest-dwelling passerine birds in the family Turdidae (thrushes), commonly known as nightingale-thrushes. The extant species are widely distributed across the Americas and are descended from a ...
'' – typical American thrushes and nightingale-thrushes (13 species) * '' Zoothera'' – Asian thrushes (21 species, including one recently extinct) * ''
Geokichla The ''Geokichla'' thrushes are medium-sized mostly insectivorous or omnivorous birds in the thrush family, Turdidae. They were traditionally listed in the '' Zoothera'', but studies suggested their placement in another genus. The genus name ...
'' – (21 species) * ''
Turdus True thrushes are medium-sized mostly insectivorous or omnivorous birds in the genus ''Turdus'' of the wider thrush family, Turdidae. The genus name ''Turdus'' is Latin for "thrush". The term " thrush" is used for many other birds of the family T ...
'' – true thrushes (88 species, including one recently extinct) See list of thrush species for more detail.


Cooking

The thrush is one of the many kinds of small bird that have in the past been trapped and eaten in much of Europe; the practice is now rare. Among traditional ways of cooking thrush were with
polenta Polenta (, ) is a dish of boiled cornmeal that was historically made from other grains. The dish comes from Italy. It may be served as a hot porridge, or it may be allowed to cool and solidify into a loaf that can be baked, fried, or grilled. ...
or grilled on a skewer, in Italy; with
juniper Junipers are coniferous trees and shrubs in the genus ''Juniperus'' () of the cypress family Cupressaceae. Depending on the taxonomy, between 50 and 67 species of junipers are widely distributed throughout the Northern Hemisphere, from the Arcti ...
berries in Belgium; and made into a
pâté ''Pâté'' ( , , ) is a paste, pie or loaf filled with a forcemeat. Common forcemeats include ground meat from pork, poultry, fish or beef; fat, vegetables, herbs, spices and either wine or brandy (often cognac or armagnac). It is often ser ...
or
terrine Terrine may refer to: * Terrine (cookware), a vessel for cooking a forcemeat loaf * Terrine (food), a forcemeat similar to pâté {{Disambiguation ...
. The French cook and cookery writer
Marie-Antoine Carême Marie Antoine (Antonin) Carême (; 8 June 178412 January 1833) was a French chef and an early practitioner and exponent of the elaborate style of cooking known as ''grande cuisine'', the "high art" of French cooking: a grandiose style of cookery ...
recommended cooking thrushes in
crépinette A crépinette is a small, flattened sausage, sometimes referred to as a sausage parcel. It is similar in shape to a sausage patty, circular, and flattened. It is made from minced or ground pork, turkey meat, turkey, veal, Lamb and mutton, lamb or ...
s and serving with sauce Périgueux.


References


External links


Thrush videos
on the Internet Bird Collection
High-resolution photo gallery of around 100 species.
{{Authority control * Taxa named by Constantine Samuel Rafinesque