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Theodosius Grigorievich Dobzhansky (russian: Феодо́сий Григо́рьевич Добржа́нский; uk, Теодо́сій Григо́рович Добржа́нський; January 25, 1900 – December 18, 1975) was a prominent Russian-American geneticist and evolutionary biologist, and a central figure in the field of evolutionary biology for his work in shaping the modern synthesis. Born in the Russian Empire, Dobzhansky emigrated to the United States in 1927, aged 27. He was a distant relation of the Russian writer
Feodor Dostoevsky Fyodor Mikhailovich Dostoevsky (, ; rus, Фёдор Михайлович Достоевский, Fyódor Mikháylovich Dostoyévskiy, p=ˈfʲɵdər mʲɪˈxajləvʲɪdʑ dəstɐˈjefskʲɪj, a=ru-Dostoevsky.ogg, links=yes; 11 November 18219 ...
. His 1937 work ''
Genetics and the Origin of Species ''Genetics and the Origin of Species'' is a 1937 book by the Ukrainian-American evolutionary biologist Theodosius Dobzhansky. It is regarded as one of the most important works of the modern synthesis, and was one of the earliest. The book populari ...
'' became a major influence on the modern synthesis. He was awarded the US
National Medal of Science The National Medal of Science is an honor bestowed by the President of the United States to individuals in science and engineering who have made important contributions to the advancement of knowledge in the fields of behavioral and social scienc ...
in 1964 and the
Franklin Medal The Franklin Medal was a science award presented from 1915 until 1997 by the Franklin Institute located in Philadelphia, Pennsylvania, U.S. It was founded in 1914 by Samuel Insull Samuel Insull (November 11, 1859 – July 16, 1938) was a Bri ...
in 1973.


Biography


Early life

Dobzhansky was born on January 25, 1900, in
Nemirov Nemyriv ( uk, Немирів, russian: Немирoв, pl, Niemirów) is a historic town in Vinnytsia Oblast (province) in Ukraine, located in the historical region of Podolia. It was the administrative center of former Nemyriv Raion (district) ...
, Russian Empire (now Nemyriv, Ukraine), the only child of Grigory Dobzhansky, a mathematics teacher, and Sophia Voinarsky. He was given a rare name,
Theodosius Theodosius ( Latinized from the Greek "Θεοδόσιος", Theodosios, "given by god") is a given name. It may take the form Teodósio, Teodosie, Teodosije etc. Theodosia is a feminine version of the name. Emperors of ancient Rome and Byzantium ...
, because he was born after his middle-aged parents prayed for a child to St. Theodosius of Chernigov. In 1910 the family moved to Kiev. At high school, Dobzhansky collected butterflies and decided to become a biologist. In 1915, he met Victor Luchnik who convinced him to specialize in
beetles Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describe ...
instead. Dobzhansky attended the
University of Kiev Kyiv University or Shevchenko University or officially the Taras Shevchenko National University of Kyiv ( uk, Київський національний університет імені Тараса Шевченка), colloquially known as KNU ...
between 1917 and 1921, where he then studied until 1924 specializing in entomology. He then moved to St Petersburg to study under
Yuri Filipchenko Yuri Aleksandrovich Filipchenko (russian: Юрий Александрович Филипченко; sometimes spelled Philipchenko) (1882 — 1930) was a Russian entomologist who coined the terms '' microevolution'' and ''macroevolution,'' as well ...
, where a '' Drosophila melanogaster'' laboratory had been established. On August 8, 1924, Dobzhansky married geneticist Natalia "Natasha" Sivertzeva, who was working with I. I. Schmalhausen in Kiev. The Dobzhanskys had one daughter, known under her married name as Sophie Coe, an anthropologist, food historian, and author, primarily known for her work on the
history of chocolate The history of chocolate began in Mesoamerica. Fermented beverages made from chocolate date back to at least 1900 BC to 1500 BC. The Mexica believed that cacao seeds were the gift of Quetzalcoatl, the god of wisdom, and the seeds once had so muc ...
. Before emigrating to the United States, Dobzhansky published 35 scientific works on entomology and genetics.


America

Dobzhansky immigrated to the United States in 1927 on a work-study scholarship from the International Education Board of the Rockefeller Foundation. Upon arriving in New York City on December 27, he joined the '' Drosophila'' Group at Columbia University working alongside Thomas Hunt Morgan and
Alfred Sturtevant Alfred Henry Sturtevant (November 21, 1891 – April 5, 1970) was an American geneticist. Sturtevant constructed the first genetic map of a chromosome in 1911. Throughout his career he worked on the organism ''Drosophila melanogaster'' with ...
. Their work provided crucial information on Drosophila cytogenetics. Additionally, Dobzhansky and his team helped establish ''
Drosophila pseudoobscura ''Drosophila'' () is a genus of flies, belonging to the family Drosophilidae, whose members are often called "small fruit flies" or (less frequently) pomace flies, vinegar flies, or wine flies, a reference to the characteristic of many species ...
'', within the genus ''Drosophila'', as a favorable model organism in evolutionary-biological studies ever since they published their influential works. Dobzhansky’s original mindset (after studying alongside
Yuri Filipchenko Yuri Aleksandrovich Filipchenko (russian: Юрий Александрович Филипченко; sometimes spelled Philipchenko) (1882 — 1930) was a Russian entomologist who coined the terms '' microevolution'' and ''macroevolution,'' as well ...
), was that there were serious doubts on using data obtained from phenomena happening in local populations ( microevolution) and phenomena happening on a global scale (
macroevolution Macroevolution usually means the evolution of large-scale structures and traits that go significantly beyond the intraspecific variation found in microevolution (including speciation). In other words, macroevolution is the evolution of taxa abov ...
).  Filipchenko also believed that there were only two types of inheritance: Mendelian inheritance of variation within species, and Non-Mendelian inheritance of variation in a macroevolutionary sense. Dobzhansky later stated that Filipchenko “bet on the wrong horse”. He followed Morgan to the
California Institute of Technology The California Institute of Technology (branded as Caltech or CIT)The university itself only spells its short form as "Caltech"; the institution considers other spellings such a"Cal Tech" and "CalTech" incorrect. The institute is also occasional ...
from 1930 to 1940. On the basis of his experiments, he articulated the idea that
reproductive isolation The mechanisms of reproductive isolation are a collection of evolutionary mechanisms, behaviors and physiological processes critical for speciation. They prevent members of different species from producing offspring, or ensure that any offspring ...
can be caused by differences in presence of microbial symbionts between populations. In 1937, he published one of the major works of the modern evolutionary synthesis, the synthesis of evolutionary biology with genetics, titled ''
Genetics and the Origin of Species ''Genetics and the Origin of Species'' is a 1937 book by the Ukrainian-American evolutionary biologist Theodosius Dobzhansky. It is regarded as one of the most important works of the modern synthesis, and was one of the earliest. The book populari ...
'', which amongst other things, defined evolution as "a change in the frequency of an allele within a
gene pool The gene pool is the set of all genes, or genetic information, in any population, usually of a particular species. Description A large gene pool indicates extensive genetic diversity, which is associated with robust populations that can surv ...
". Dobzhansky's work was instrumental in spreading the idea that it is through mutations in genes that natural selection takes place. Also in 1937, he became a
naturalized citizen Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the in ...
of the United States. During this time, he had a very public falling out with one of his ''Drosophila'' collaborators,
Alfred Sturtevant Alfred Henry Sturtevant (November 21, 1891 – April 5, 1970) was an American geneticist. Sturtevant constructed the first genetic map of a chromosome in 1911. Throughout his career he worked on the organism ''Drosophila melanogaster'' with ...
, based primarily in professional competition. He returned to Columbia University from 1940 to 1962. Among his students was geneticist Bruce Wallace. In 1941, Dobzhansky was awarded the Daniel Giraud Elliot Medal from the National Academy of Sciences. In 1943, the
University of São Paulo The University of São Paulo ( pt, Universidade de São Paulo, USP) is a public university in the Brazilian state of São Paulo. It is the largest Brazilian public university and the country's most prestigious educational institution, the best ...
awarded him an honorary doctorate. He was one of the signatories of the 1950 UNESCO statement ''
The Race Question The Race Question is the first of four UNESCO statements about issues of race. It was issued on 18 July 1950 following World War II and Nazi racism to clarify what was scientifically known about race, and as a moral condemnation of racism.< ...
''. He then moved to the Rockefeller Institute (shortly to become Rockefeller University) until his retirement in 1971. In 1972 he was elected the founding president of the
Behavior Genetics Association The Behavior Genetics Association (BGA) is a learned society established in 1970 and which promotes research into the connections between heredity and behavior, both human and animal. Its members support education and training in behavior genetics ...
, and was recognized by the society for his role in behavior genetics, and the founding of the society by the creation of the Dobzhansky Award (for a lifetime of outstanding scholarship in behavior genetics). Dobzhansky’s work in the field of evolutionary genetics, with the help of Sewall Wright, integrated standards of the theoretical, natural historical, and experimental work. Dobzhansky was elected a Foreign Member of the Royal Society (ForMemRS) in 1965. In 1970, he published ''Genetics of the evolutionary process''. Dobzhansky was renowned as the president of the Genetics Society of America in 1941, president of the
American Society of Naturalists The American Society of Naturalists was founded in 1883 and is one of the oldest professional societies dedicated to the biological sciences in North America. The purpose of the Society is "to advance and diffuse knowledge of organic evolution and o ...
in 1950, president of the
Society for the Study of Evolution The Society for the Study of Evolution is a professional organization of evolutionary biologists. It was formed in the United States in 1946 to promote evolution and the integration of various fields of science concerned with evolution and to organ ...
in 1951, president of the
American Society of Zoologists The Society for Integrative and Comparative Biology is organized to integrate the many fields of specialization which occur in the broad field of biology.. The society was formed in 1902 as the American Society of Zoologists, through the merger of ...
in 1963, a member of the Board of Directors of the
American Eugenics Society American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...
in 1964, and president of the American
Teilhard de Chardin Pierre Teilhard de Chardin ( (); 1 May 1881 – 10 April 1955) was a French Jesuit priest, scientist, paleontologist, theologian, philosopher and teacher. He was Darwinian in outlook and the author of several influential theological and philo ...
Association in 1969. Dobzhansky’s research and studies allowed him to travel the world and receive honorary degrees in Australia, Belgium, Brazil, Canada, Denmark, England, Germany, Italy, Japan, and Sweden.


''Genetics and the Origin of Species''

Theodosius Dobzhansky published three editions of his book ''
Genetics and the Origin of Species ''Genetics and the Origin of Species'' is a 1937 book by the Ukrainian-American evolutionary biologist Theodosius Dobzhansky. It is regarded as one of the most important works of the modern synthesis, and was one of the earliest. The book populari ...
''. Although the book was meant for people with a background in biology, it was easily understood. In the fields of genetics and evolution, Dobzhansky’s book is acknowledged as one of the most important books ever written. With each revision of ''Genetics and the Origin of Species'', Dobzhansky added new material on crucial, up to date topics, and removed material he deemed to be no longer crucial. His book sparked trends in genetic research and theory. The first edition of ''Genetics and the Origin of Species'' (1937) highlighted the most recent discoveries in genetics and how they applied to the concept of evolution. The book starts by addressing the problem of evolution and how modern discoveries in genetics could help find a solution. The book covers the chromosomal basis of Mendelian Inheritance, how the effects from changes in chromosomes greater than gene mutations are common and acceptable, and how mutations form racial and specific differences. Dobzhansky explained how three levels could describe the processes of evolutionary population genetics: (1) the origin of raw materials by mutations of genes and chromosomes, (2) the changes in populations by changes in frequencies and combinations of mutations, (3) the fixation of changes by reproductive isolation. To support his writing and research, the bibliography was twenty-eight pages long with around six hundred sources. In Dobzhansky’s second edition of ''Genetics and the Origin of Species'' (1941), four years had gone by and he was able to add more research and advancements made in genetics. Around half of the new research he found was added to the last two chapters in his book: Patterns of Evolution, and Species as Natural Units. In the second to last chapter, Patterns of Evolution, Dobzhansky explained how on the path to a new adaptation, a method could be used to where a species could go through a less adaptive stage. The last chapter, Species as Natural Units, Dobzhansky explained some of the contributions made in genetics to what was called “the new systematics.” Dobzhansky’s second edition of the book also had twice as many sources in the bibliography than the first edition. In the third revision of ''Genetics and the Origin of Species'' (1951), Dobzhansky rewrote all ten chapters on: Isolating Mechanisms, Mutation in Populations, Organic Diversity, Heredity and Mutation, Race Formation, Selection, Adaptive Polymorphism, Hybrid Sterility, Species as Natural Units, and Patterns of Evolution. Dobzhansky decided to remove the chapter on Polyploidy in the third edition. The new chapter on Adaptive Polymorphism highlighted Dobzhansky’s research since the second edition. He included precise, quantitative evidence on effective natural selection in laboratory and free populations.


Debate about race

Theodosius Dobzhansky and
Ashley Montagu Montague Francis Ashley-Montagu (June 28, 1905November 26, 1999) — born Israel Ehrenberg — was a British-American anthropologist who popularized the study of topics such as race and gender and their relation to politics and development. He w ...
debated the use and validity of the term "race" over a period of many years without reaching an agreement. The debate has continued. Montagu argued that "race" was so laden with toxic associations that it was a word best eliminated from science completely. Dobzhansky strongly disagreed. He argued that science should not give in to the misuses to which it had been subjected. The two men never reached an agreement, which led Dobzhansky to say in 1961, while commenting on Montagu's autobiography, "The chapter on 'Ethnic group and race' is, of course, deplorable, but let us say that it is good that in a democratic country any opinion, no matter how deplorable, can be published" (Farber 2015 p. 3). The concept of "race" has been important in many life science disciplines; the modern synthesis revolutionized the concept of race, moving it from a strictly morphological definition based on "racial types" in humans, to a definition focused on populations differing in gene frequencies. This was done in hopes that its foundation in population genetics would undermine the deeply ingrained social prejudices associated with "race". Dobzhansky was confident that mixing races created no serious medical issues. Dobzhansky's experience with breeding fruit flies came into play when he made this conclusion. The only medical issue Dobzhansky found in this breeding was when certain crosses could lead to having infertile offspring. However, Dobzhansky noticed no such problems when humans from different populations reproduced. When anthropologists at the time were trying to compare the means of physical measurements of people from different races Dobzhansky argued that these means had no value because there was more variation between the individuals of each population than there was among the groups (Farber 2011 p. 63). However, Dobzhansky's work and beliefs on genetics and evolution created opposition with his views on race mixing. First, that race has to do with groups and not individuals and so in this instance it is not races that mix, it is individuals. Second, if races do not mix then they will become different species, so therefore they have to mix. All of the races that currently exist are products of past mixed races, so according to Dobzhansky there is no pure race. Third, when race had been discussed in the past it was all about comparing means of trait to which this made no sense to Dobzhansky (Farber 2011 p. 65-67). His concern with the interface between humans and biology may have come from different factors. The main factor would be the race prejudice that contributed in Europe that triggered
WWII World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the vast majority of the world's countries—including all of the great powers—forming two opposing ...
. His concern also dealt with religion in human life which he speaks about in his book ''The Biology of Ultimate Concern'' in 1967. "The pervasiveness of genetic variation provides the biological foundation of human individuality". Dobzhansky talks about in great detail that "human nature has 2 dimensions: the biological, which mankind shares with the rest of life, and the cultural, which is exclusive to humans." Both of these are believed to have come from "biological evolution and cultural evolution". Dobzhansky sought to put an end to the pseudoscience that purports genetic makeup to determine race, and thus rank in society. Harrison E. Salisbury wrote in a New York Times review of Dobzhansky's book ''Heredity and the Future of Man'' that Dobzhansky could not, together with other scientists, agree upon what defines a race. Dobzhansky stated that a true bloodline for man could not be identified. He did not believe that a person's genetic makeup decided whether or not he would be a great man but rather that man "has the rare opportunity 'to direct his evolution'".


Final illness and the "Light of Evolution"

Dobzhansky's wife Natasha died of
coronary thrombosis Coronary thrombosis is defined as the formation of a blood clot inside a blood vessel of the heart. This blood clot may then restrict blood flow within the heart, leading to heart tissue damage, or a myocardial infarction, also known as a heart at ...
on February 22, 1969. Earlier (on June 1, 1968), Theodosius had been diagnosed with
lymphocytic leukemia Lymphoid leukemias are a group of leukemias affecting circulating lymphocytes, a type of white blood cell. The lymphocytic leukemias are closely related to lymphomas of the lymphocytes, to the point that some of them are unitary disease entities t ...
(a chronic form of leukemia), and had been given a few months to a few years to live. He retired in 1971, moving to the
University of California, Davis The University of California, Davis (UC Davis, UCD, or Davis) is a public land-grant research university near Davis, California. Named a Public Ivy, it is the northernmost of the ten campuses of the University of California system. The institu ...
where his student Francisco J. Ayala had been made assistant professor, and where he continued working as an emeritus professor. He published one of his most famous essays "
Nothing in Biology Makes Sense Except in the Light of Evolution "Nothing in Biology Makes Sense Except in the Light of Evolution" is a 1973 essay by the evolutionary biologist Theodosius Dobzhansky, criticising anti-evolution creationism and espousing theistic evolution. The essay was first published in '' ...
" in 1973, influenced by the paleontologist and priest Pierre Teilhard de Chardin. By 1975, his leukemia had become more severe, and on November 11 he traveled to San Jacinto, California, for treatment and care. Working till his last day as a Professor of Genetics, Dobzhansky died (from heart failure) on December 18, 1975, in Davis, California. He was cremated, and his ashes were scattered in the Californian wilderness.


Religious beliefs

Ernst Mayr stated: "On the other hand, famous evolutionists such as Dobzhansky were firm believers in a personal God." Dobzhansky himself spoke of God as creating through evolution, and was a communicant of the Eastern Orthodox Church. Although Dobzhansky came from a long line of Eastern Orthodox priests, later in life, he had doubts about a conventional afterlife. He stated that if a
Heaven Heaven or the heavens, is a common religious cosmological or transcendent supernatural place where beings such as deities, angels, souls, saints, or venerated ancestors are said to originate, be enthroned, or reside. According to the belie ...
did exist, it would not be a place where one could find all the answers about life in an instant. It would rather be a place where performing experiments would give rise to precise and explicit results.


Publications

During his career, Dobzhansky published widely in books and peer reviewed
scientific journal In academic publishing, a scientific journal is a periodical publication intended to further the progress of science, usually by reporting new research. Content Articles in scientific journals are mostly written by active scientists such as ...
s:


Books

* Sinnott, E.W., Dunn, L.C and Dobzhansky, Th. 1925. ''Principles of Genetics''. McGraw-Hill. (5 editions: 1925, 1932, 1939, 1950, 1958; Dobzhansky co-editor only on 1950 & 1958 editions). * Dobzhansky, Th. 1937. ''
Genetics and the Origin of Species ''Genetics and the Origin of Species'' is a 1937 book by the Ukrainian-American evolutionary biologist Theodosius Dobzhansky. It is regarded as one of the most important works of the modern synthesis, and was one of the earliest. The book populari ...
''. Columbia University Press, New York. (2nd ed., 1941; 3rd ed., 1951) * ''The Biological Basis of Human Freedom'' (1954). * Dunn, L. C., & Dobzhansky, Th. 1946. ''Heredity, Race, and Society''. The New American Library of World Literature, Inc., New York. * Dobzhansky, Th. 1955. ''Evolution, Genetics, & Man''. Wiley & Sons, New York. * Dobzhansky, Th. 1962. ''Mankind Evolving''. Yale University Press, New Haven, Connecticut. * Dobzhansky, Th. 1966. ''Heredity and the Nature of Man''. Harcourt, Brace & World Inc., New York, New York. * Dobzhansky, Th. 1967. ''The Biology of Ultimate Concern''. New American Library, New York. * Dobzhansky, Th. 1970. ''Genetics of the Evolutionary Process''. Columbia University Press, New York. * Dobzhansky, Th. 1973. ''Genetic Diversity and Human Equality''. Basic Books, New York. * Dobzhansky, Th., F.J. Ayala, G.L. Stebbins & J.W. Valentine. 1977. ''Evolution''. W.H. Freeman, San Francisco. * Dobzhansky, Th. 1981. ''Dobzhansky's Genetics of Natural Populations I-XLIII''. R.C. Lewontin, J.A. Moore, W.B. Provine & B. Wallace, eds. Columbia University Press, New York. (reprints the 43 papers in this series, all but two of which were authored or co-authored by Dobzhansky) * Dobzhansky, Th., & Boesiger, E. 1983. ''Human Culture, A Moment in Evolution''. Columbia University Press, New York.


Papers

* *


Recensions

* Dobzhansky, Th. Wrote a recension of "The origin of races" by the anthropologist
Carleton S. Coon Carleton Stevens Coon (June 23, 1904 – June 3, 1981) was an American anthropologist. A professor of anthropology at the University of Pennsylvania, lecturer and professor at Harvard University, he was president of the American Association of ...
. Dobzhansky rejected Coon's theory of independent origin of identical mutations, but he did agree that selection favored a sapiens-like genotype in all proto-human populations, and expressed the theory that all sapiens-alleles existed at a low frequency in all erectus-populations, and that the statistical composition of the gene pool shifted from erectus to sapiens in multiple populations independently.


References


External links


Colloquium on ''Genetics and the Origin of Species''
with
biography



Theodosius Dobzhansky: A Man For All Seasons
by Francisco J. Ayala
The Theodosius Dobzhansky Papers
American Philosophical Society The American Philosophical Society (APS), founded in 1743 in Philadelphia, is a scholarly organization that promotes knowledge in the sciences and humanities through research, professional meetings, publications, library resources, and communit ...

National Academy of Sciences Biographical Memoir
* {{DEFAULTSORT:Dobzhansky, Theodosius 1900 births 1975 deaths People from Nemyriv People from Bratslavsky Uyezd American geneticists Ukrainian geneticists American entomologists Ukrainian entomologists Evolutionary biologists Soviet emigrants to the United States Modern synthesis (20th century) National Medal of Science laureates Columbia University alumni Columbia University faculty Critics of creationism Critics of Lamarckism Foreign Members of the Royal Society Rockefeller University faculty Theistic evolutionists 20th-century Ukrainian zoologists Eastern Orthodox Christians from Ukraine Eastern Orthodox Christians from the United States Soviet entomologists Soviet geneticists Members of the Royal Swedish Academy of Sciences